1 
The Development of the Gametophytes and 
Embryogeny in Cunninghamia sinensis 1 ). 
By 
Kiichi Miyake. 
With Plates I—V and 2 Figures in the Text. 
Reeent embryological studies of Arnoldi, Coker and Lawson 
have shown quite conclusiyely that the family Taxodieae is not at 
all a natural group, and that the present Classification of the Co¬ 
rd ferne, so f'ar as this group is concerned, should be entirely re- 
arranged. Amol di (1900) has already suggested that, of the 
genera of this family, Taxodium, Cryptomeria and perhaps also 
Cunninghamia 2 ), should be united with the Cupressineae and that 
Sequoia and Sdadopitys should each constitute a family by itself. 
This Suggestion, being followed by Coker (1903) and Lawson 
(1904), seems to be well founded. But before making any de¬ 
finite new arrangement of the group, it is very desirable that the 
life-history of all the genera should be thoroughly worked out. 
Of eight genera 3 ) belonging to the Taxodieae, we have only 
the fairly complete embryological records of three, namely those 
of Sequoia ; (Shaw, 1896; Lawson, 1904 a), Taxodium (Coker, 
1903) and Cryptomeria (Lawson, 1904b). Our knowledge about 
the life-history of the genus Cunninghamia is very meager, being 
based entirely on the fragmentary records of Am old i (1900) who 
has studied the genus incidentally, with insufficient material, in 
his comparative investigations of the Taxodieae. The present work 
was begun in the summer of 1905 with the hope of filling up the 
q A preliminary note was published in Bot. Mag. Tokyo. Vol. 22. 
(Miyake, 1908.) 
2 ) He was not quite sure ab out Cunninghamia as the following statement 
shows: „Es wird also nach meiner Meinung die von Eichler als Pinoideae- 
Abietineae-Taxodiinae bezeichnete Gruppe in drei Familien geteilt. Die 
erste Familie Sequoiaceae — enthält zwei Gattungen: Sequoia and Welling- 
tonia; Taxodium und Cryptomeria , vielleicht auch Cunninghamia (?), werden 
mit den Cupressineen vereinigt; Sdadopitys wird am besten als Repräsentant 
einer eigenen Familie Sciadopiteae — bezeichnet.” (Arnoldi, 1900, p. 23.) 
3 ) Seven genera have hitherto been included in the Taxodieae, and an 
another genus Taiwania was recently established by Hayata (1906) basing 
on a new Conifer from Formosa. 
Beihefte Bot. Centralbl. Bd. XXVII. Abt. I. Heft 1. 
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