298 
Kusano, Further Studies on Aeghietia imlica. 
In Aeginetia the formation of the hair-tendrils is alone sufficient 
to avoid sucli a danger. 
In tlie second place, Aeginetia shows some transitional States 
between autopbytic and the most advanced parasitic life. In most 
hemiparasites, or more strictly speaking, green parasites such as 
Santalaceae (Kusano, 1906, Barber 1906, 1907), Ehinanthaceae 
(Heinricher, 1901, 1902) and Loranthaceae, the germination is 
neither associated with the presence of the host, nor have they 
any marked tendency to select their host. But some holoparasites 
or at last Orobanchaceae, hitherto stndied, have acquired thehabit 
of not developing and even of not germinating without the presence 
of the roots of their proper host. While thus the intimate relation 
of the parasite and host-root is in this case restricted to certain 
limited species of plants, Aeginetia shows itself to be many-sided 
in this respect: In Orobanche and Latkraea the selection of the 
host takes place already at the period of germination, it takes 
place in Aeginetia at a later period. Thus certain variations ob- 
served to occur in the Orobanchaceae in their behaviour towards 
the host-roots a study of other species of the same family is very 
desirable. 
The chief results of the experiments described in the fore- 
going pages may be summarised as foUows: 
1. The germination of Aeginetia- does not take place in 
water, moist chamber or soil. It requires always the Stimulus of 
the root of other plants. 
2. The seed kept dry for two years loses its germinating power. 
3. The plants which stimnlate the seed to germination may 
be Vascular Cryptogams, Gymnosperms, or Angiosperms. 
4. The stimulant is an unknown substance that is perhaps 
excreted by active roots of all higher plants. 
5. The development of the seedlings takes place only on 
certain species of Monocotyledons. Its conditions are entirely 
different from those that are necessary for the germination of the 
seeds, the former being fulfiUed only by certain plants while the 
latter are found in the roots of all higher plants. 
6. The first change that takes place during the germination 
is the sweUing of the epidermal cells at the radicular end of the 
embryo and their transformation into the hair-tendrils. 
7. The seedlings are much reduced in form, and before they 
are connected with the host no mnltiplication of cells take place. 
8. The seedlings develop, when attached to the host, spherical 
tnbercles. They are formed by the meristematic tissne under the 
hair-tendrils. 
9. For the mnltiplication of cells in the seedlings certain 
Stimulus from the host-roots to which the hair-tendrils are sensitive 
seems to be reqnired. 
10. The tnbercles become differentiated first into the primary 
haustorium at the frontal portion, and then into the shoot and 
root-syStern at the other portions. 
Botanical Institute, Agricultural College, Tokyo. 
June, 1908. 
