453 
The Systematic Anatomy of the 
genus Canephora. 
By 
H. F. Wernham, B. Sc. 
Assistant in the Department of Botany, British Museum; 
Lecturer in Botany at the Goldsmiths’ College, University of London. 
With 7 figures in the text. 
The examination from the systematic standpoint of the spe- 
cimens of Canephora in the National and Kew Herbaria 1 ) has led 
me to investigate the anatomy of these curious plants as fully as 
may have been consistent with the scanty and somewhat poor 
material available, which was taken from herbarium specimens. 
The bulk of the tissues, being more or less lignified, is tolerably 
well preserved; but the softer portions, particularly the phloem, 
have perished to a considerable extent, 
This rare genus of Rabiaceae is, so far as onr present know- 
ledge extends, confined to Madagascar. 
The five species hitherto described all reveal more or less 
strongly marked xerophytic characters; they are shrubs or trees, 
with leathery leaves, and destitute of indumentum. The rather 
small campanulate ilowers are borne two or three together in the 
hollowed apices of axillary fiattened branches (Fig. 1). The corolla 
is contorted in aestivation, and the bilocular ovary contains a few 
ovules, maturing into a few-seeded berry, so that the genus finds 
a place in the section Cinchonoideae-Gardenihiae-Gardenieae . 2 ) 
We may proceed forthwith to such anatomical details as have 
been ascertained in the course of the enquiry. Three only of the 
five known species have been available for the purpose of this 
investigation — C. madagascariensis Gmelin, upon which the genus 
was founded; C. angnstifolia and C. Goudotii , described by myself 
in the Journal of Botany (loc. cit.). 
l ) Wernham, Journ. of Botany. Vol. XLIX. 1911. p. 77—82. 
*) Schumann, K., u. Engler, A., Xat. Pflanzenfam. IV. 4. p. 80. 
