Wernham, The Systematic Anatomy of tke genus Canephora. 
467 
C. Goudotii Wernham. 
I. Stern. The pith is relatively much bulkier than in the 
other two species. The cells are roughly circular in section, and 
are thick-walled throughout; no lacunae appear, and this may in- 
dicate the absence of thin-walled tissue in the pith of the living 
plant, in contrast with the species dealt with above. Not much 
tannin is present in the pith; calcium oxalate is fairly abundant, 
a good deal of it appearing as crystal-sand, but mostly, perhaps, 
in the form of small clnster crystals. 
The stem xylem is essentially similar to that of the other 
two species, but the primary wood is more sharply distinguished 
from the pith, its elements being considerably smaller. The fibres 
are very thick-walled and the yessels smaller in section. The 
wood-ring is perforated by a good many medullary rays, of the 
type described for C. angustifolia\ there is no other parenchyma 
associated with the xylem, but a few small lacunae are to be seen 
in the region of the primary xylem. 
The phloem consists of a narrow band almost obliterated 
with tannin, and containing also an abundance of crystal-sand. 
Immediately surrounding this is a fairly continuous fibrous sheath, 
as in the other species; this comprises only a single layer for the 
greater part of its extent. 
The cortex is composed mostly of thick-walled cells. Tannin 
is not so conspicuous a feature of the celhcontents as in the other 
species, but calcium oxalate, in the form of crystal-sand and small 
cluster crystals, is very abundant. This is contained mostly in 
cells which appear to be larger than their neighbours, and in some 
cases definite sacs, formed apparently from two or three cells, are 
receptacles for an abundance of calcium oxalate. 
The epidermis displays no distinctive features; no hypodermis 
appears to be differentiated. 
II. Leaf. a) Petiole. This, unlike the two already described, 
is approximately circular in section (Fig. 3, c). The vascular System 
consists, as in the previous species, of one large median bündle 
and two lateral small ones. The extremities of the forrner are 
involute, and are separated by a relatively broad gap. 
The xylem of the median bündle is dense, consisting entirely 
of vessels, which are traversed by narrow medullary rays infiltrated 
with tannin. The phloem appears to be composed almost entirely 
of sieve-tube tissue, appearing as a two to four-layered band ex- 
tending continuously round the wood ring as far as the edges of 
the gap; this tissue is packed with crystal-sand of calcium oxalate. 
At several points the phloem “invades“ the xylem. 
As in C. angustifolia, no definite fibrous sheath surrounds 
the phloem; a few large fibres, isolated for the most part, appear 
in the neighbourhood of the gap. 
The gap is in reality but one cell-layer broad, its relatively 
great width being due to the large size of the lumina of the cells 
composing this layer. The tissue in the concavity of the bündle 
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