472 
Wernham, The Systematic Anatomy of the genus Canephora. 
parallel to the pore and so the stomata conform to the 
so-called “Rubiaceous“ type. 1 ) 
Thepeduncle of the inflorescence is invariablyborne 
in the axil of a leaf (see fig. 1), and so must be regarded 
morphologically as a stem-structure; this estimate of its 
nature is borne out by the fact that the main bündle is a 
completely-closed cylinder. The faint Suggestion of dorsi- 
ventrality, both internally and externally, is curious 
(supra, p. 463). 
As regards the cell-contents, tannin is plentiful, 
especially in the exterior layers (excepting, of course, 
the epidermis). It abounds in the phloem of all parts, and 
occurs thijoughout the ground-pa'renchyma, pith, andintra- 
stelar tissue generally, and frequently in the medullary 
rays. In the leaf, tannin is especially abundant in the 
palisade tissue. 
Calcium oxalate occurs in the form either of crystal- 
sand or of cluster-crystals; theformer, moreover, displays 
a marked tendency to become aggregated into amorphous 
masses. No raphides were observed. Crystal-sand is pre¬ 
sent generally in the ground tissue, both intra- andextra- 
stelar (pith, cortex etc.); it is usually yery abundant in 
the phloem. 
The foregoing research has been conducted in the Botanical 
Department of Goldsmiths College, University of London; and my 
acknowledgements are due to the Trustees of the British Museum 
for the use of the material, which was taken from the National 
Herbarium. I have also to thank Mr. Kench, of the Botanical De¬ 
partment, British Museum, for figure 1, which he copied from 
Drake’s Hist. Nat. des Plantes in the Hist. Phys. Nat. et Pol. de 
Madagascar of Grandidier (vol. XXXVI, tom. VI, pl. 444, fig. 1). 
b V. Solereder, Systematic Anatomy of the Dicotyledons. Transl. Oxford, 
(Boodle and Fritsch) 1908. II. p. 1078. 
