U.S. Department 
of Commerce 
Wilbur L. Ross Jr. 
Secretary 
National Oceanic 
and Atmospheric 
Administration 
Neil A. Jacobs 
Acting Administrator 
National Marine 
Fisheries Service 
Chris Oliver 
Assistant Administrator 
for Fisheries 
The Fishery Bulletin (ISSN 0090-0656) 
is published quarterly by the Scientific 
Publications Office, National Marine 
Fisheries Service, NOAA, 7600 Sand 
Point Way NE, Seattle, WA 98115-0070. 
Although the contents of this publica¬ 
tion have not been copyrighted and may 
be reprinted entirely, reference to source 
is appreciated. 
The Secretary of Commerce has deter¬ 
mined that the publication of this peri¬ 
odical is necessary according to law for 
the transaction of public business of this 
Department. Use of funds for printing of 
this periodical has been approved by the 
Director of the Office of Management and 
Budget. 
For Sale by the Superintendent of 
Documents, U.S. Government Printing 
Office, Washington, DC 20402. Subscrip¬ 
tion price per year: $32.00 domestic and 
$44.80 foreign. Cost per single issue: 
$19.00 domestic and $26.60 foreign. See 
back for order form. 
Scientific Editor 
Jose I. Castro 
National Marine Fisheries Service, NOAA 
263 13th Avenue South 
Saint Petersburg, Florida 33701 
Managing Editor 
Kathryn Dennis 
National Marine Fisheries Service, NOAA 
Scientific Publications Office 
1845 Wasp Blvd., Bldg. 176 
Honolulu, Hawaii 96818 
Associate Editor 
Cara Mayo 
National Marine Fisheries Service, NOAA 
Scientific Publications Office 
7600 Sand Point Way NE 
Seattle, Washington 981 15-0070 
Sharyn Matriotti (retired*) 
National Marine Fisheries Service, NOAA 
Scientific Publications Office 
7600 Sand Point Way NE 
Seattle, Washington 98115-0070 
Editorial Board 
Henry L. Bart Jr. 
Shannon L. Cass-Calay 
William B. Driggers III 
Gretchen L. Grammer 
Richard Langton 
Richard S. McBride 
Richard L. Merrick 
Richard D. Methot 
Lisa J. Natanson 
Joseph M. Quattro 
Mark S. Peterson 
John F. Walter III 
Bruce Wright 
Tulane University, New Orleans, Louisiana 
National Marine Fisheries Service, Miami, Florida 
National Marine Fisheries Service, Pascagoula, Mississippi 
South Australian Research and Development Institute, Adelaide, Australia 
National Marine Fisheries Service, Walpole, Maine (retired) 
National Marine Fisheries Service, Woods Hole, Massachusetts 
National Marine Fisheries Service, Falmouth, Massachusetts (retired) 
National Marine Fisheries Service, Seattle, Washington 
National Marine Fisheries Service, Narragansett, Rhode Island 
University of South Carolina, Columbia, South Carolina 
University of Southern Mississippi, Ocean Springs, Mississippi 
National Marine Fisheries Service, Miami, Florida 
Aleutian Pribilof Island Association, Anchorage, Alaska 
*Sharyn Matriotti, a superb and dedicated editor, retired in December 2018 after 29 years of service 
as managing editor of Fishery Bulletin. 
Fishery Bulletin website: https://spo.nmfs.noaa.gov/fb.htm 
The Fishery Bulletin carries original research reports on investigations in fishery sci¬ 
ence, engineering, and economics. It began as the Bulletin of the United States Fish Commis¬ 
sion in 1881; it became the Bulletin of the Bureau of Fisheries in 1904 and the Fishery Bulle¬ 
tin of the Fish and Wildlife Service in 1941. Separates were issued as documents through vol¬ 
ume 46; the last document was no. 1103. Beginning with volume 47 in 1931 and continuing 
through volume 62 in 1963, each separate appeared as a numbered bulletin. A new system 
began in 1963 with volume 63 in which papers are bound together in a single issue. Beginning 
with volume 70, number 1, January 1972, Fishery Bulletin became a periodical, issued quarterly. In 
this form, it is available by subscription from the Superintendent of Documents, U.S. Government 
Printing Office, Washington, DC 20402. It is also available free in limited numbers to libraries, 
research institutions, state and federal agencies, and in exchange for other scientific publications. 
