330 
Fishery Bulletin 117(4) 
A Female 
. , ■ 150 
o Male 
. _ Female 100 
— Male 
28 32 
o Unfished 
A Fished 
■ — Unfished 
— Fished 
ii i-1-1-1-1 
8 12 16 20 24 28 32 
500 
A ^ 
0 4 
8 12 16 20 24 28 32 
o Unfished 
A Fished 
-Unfished 
- Fished 
Age (years) 
8 12 16 20 24 28 32 
Figure 4 
Lengths at age and fitted growth curves from the von Bertalanffy growth function for goldeneye 
jobfish (Pristipomoides flavipinnis ) collected in the Samoa Archipelago in 2012 and 2016, by 
(A) sex (female and male) and (B) fishing pressure (fished and unfished areas), and for goldflag job- 
fish (P. auricilla) collected in 2014 in the Mariana Archipelago, by (C) sex and (D) fishing pressure. 
fish with the most common ages (4—6% for fish 3-6 years 
old). Goldflag jobfish had a growth pattern between fished 
and unfished areas that was the opposite of that observed 
for goldeneye jobfish in the Samoa Archipelago. They grew 
more slowly (0.06 difference in K) and reached a larger 
(difference of 33.4 mm FL) in the unfished areas than in 
the fished areas (Fig. 4, Table 1). These differences in 
growth of goldflag jobfish between fished and unfished 
areas were further confirmed by 2-10% differences in the 
EVB-predicted FL at age for fish with the most common 
ages (3-6 years). Overall, the fitted growth curve indicates 
that growth was similar to that of goldeneye jobfish; it 
reaches an asymptote between ages 6 and 9 or between 
20% and 30% of the maximum age (Fig. 4). 
Mortality 
The multinomial catch curves provide a good fit to the 
observed age compositions for goldeneye jobfish and gold- 
flag jobfish, as evident in the predicted proportions at age 
(Fig. 5). The estimated values of total mortality from the 
unfished areas, assumed to be an estimate of natural mor¬ 
tality, were 0.22 (95% Cl: 0.19-0.25) for goldeneye jobfish 
and 0.18 (95% Cl: 0.15-0.21) for goldflag jobfish (Table 3). 
Use of Hoenig’s (1983) updated natural mortality estimator 
(Then et al., 2015) produced comparable natural mortality 
estimates for these same unfished areas (goldeneye job¬ 
fish: 0.23; goldflag jobfish: 0.20), whereas use of the Pauly 
estimator (Then et al., 2015) produced lower estimates 
(goldeneye jobfish: 0.17; goldflag jobfish: 0.18). Using the 
maximum age natural mortality estimator but substituting 
the maximum age from the fished areas produced greater 
estimates of natural mortality for goldeneye jobfish (0.59; 
maximum age: 10) and goldflag jobfish (0.35; maximum 
age: 18). The values of total mortality from the fished areas, 
calculated by using the multinomial catch curve, are 0.89 
(95% Cl: 0.36-1.42) for goldeneye jobfish and 0.42 (95% Cl: 
0.34—0.50) for goldflag jobfish. The exploitation rates ( E ), 
calculated as fishing mortality divided by total mortality, 
for both species differed greatly depending on whether nat¬ 
ural mortality was determined by using the maximum age 
from the fished or the unfished areas. Using the natural 
