302 
Fishery Bulletin 117(4) 
Gertseva, V., S. E. Matson, and J. Cope. 
2017. Spatial growth variability in marine fish: example from 
Northeast Pacific groundfish. ICES J. Mar. Sci. 74:602-1613. 
Haltuch, M. A., and A. Hicks. 
2009. Status of the U.S. petrale sole resource in 2008, 148 p. 
Pac. Fish. Manage. Counc., Portland, OR. [Available from 
website.] 
Haltuch, M. A., A. Hicks, and K. See. 
2011. Status of the U.S. petrale sole resource in 2010, 175 p. 
Pac. Fish. Manage. Counc., Portland, OR. [Available from 
website.] 
Haltuch, M. A., K. Ono, and J. Valero. 
2013. Status of the U.S. petrale sole resource in 2012, 234 p. 
Pac. Fish. Manage. Counc., Portland, OR. [Available from 
website.] 
Hannah, R. W., S. J. Parker, and E. L. Fruh. 
2002. Length and age at maturity of female petrale sole 
(Eopsetta jordani ) determined from samples collected 
prior to spawning aggregations. Fish. Bull. 100:711-719. 
Harry, G. Y., Jr. 
1959. Time of spawning, length at maturity, and fecundity of 
the English, petrale, and Dover soles ( Parophrys vetulus, 
Eopsetta jordani, and Microstomus pacificus, respectively). 
Fish. Comm. Oregon., Res. Briefs 7(1):5—13. 
Hixon, M. A., D. W. Johnson, and S. M. Sogard. 
2014. BOFFFFs: on the importance of conserving old-growth 
age structure in fishery populations. ICES J. Mar. Sci. 
71:2171-2185. 
Humason, G. L. 
1972. Animal tissue techniques, 3 rd ed., 468 p. W. H. Freeman, 
San Francisco, CA. 
Keller, A. A., J. R. Wallace, B. H. Horness, O. S. Hamel, and 
I. J. Stewart. 
2012. Variations in eastern North Pacific demersal fish bio¬ 
mass based on the U.S. west coast groundfish bottom trawl 
survey (2003-2010). Fish. Bull. 110:205-222. 
Ketchen, K. S., and F. C. R. Forrester. 
1966. Population dynamics of the petrale sole, Eopsetta jor¬ 
dani, in waters off western Canada. Fish. Res. Board Can., 
Bull. 153, 204 p. 
Kruse, G. H., and A. V. Tyler. 
1983. Simulation of temperature and upwelling effects on the 
English sole ( Parophrys vetulus) spawning season. Can. 
J. Fish. Aquat. Sci. 40:230-237. 
Lowerre-Barbieri, S. K., K. Ganias, F. Saborido-Rey, H. Murua, 
and J. R. Hunter. 
2011. Reproductive timing in marine fishes: variability, tempo¬ 
ral scales, and methods. Mar. Coast. Fish. 3:71-91. 
McBride, R. S., M. J. Wuenschel, P. Nitscheke, G. Thorton, and 
J. R. King. 
2013. Latitudinal and stock-specific variation in size- and 
age-at-maturity of female winter flounder, Pseudopleu- 
ronectes americanus, as determined with gonad histology. 
J. Sea Res. 75:41-51. 
McElroy, W. D., M. J. Wuenschel, Y. K. Press, E. K. Towle, and 
R. S. McBride. 
2013. Differences in female individual reproductive potential 
among three stocks of winter flounder, Pseudopleuronectes 
americanus. J. Sea Res. 75:52-61. 
Murua, H., and F. Saborido-Rey. 
2003. Female reproductive strategies of marine fish species 
of the North Atlantic. J. Northwest Atl. Fish. Sci. 3:23-31. 
Parrish, R. H., C. S. Nelson, and A. Bakun. 
1981. Transport mechanisms and reproductive success of 
fishes in the California Current. Biol. Oceanogr. 1:175-203. 
Pedersen, M. G. 
1975. Movement and growth of petrale sole (Eopsetta jordani) 
tagged off Washington and southwest Vancouver Island. J. 
Fish. Res. Board Can. 32:2169-2177. 
Porter, P. 
1964. Notes on fecundity, spawning, and early life history of 
petrale sole (Eopsetta jordani), with descriptions of flat¬ 
fish larvae collected in the Pacific Ocean off Humboldt 
Bay, California. M.S. thesis, 98 p. Humboldt State Coll., 
Areata, CA. 
R Core Team. 
2017. R: a language and environment for statistical com¬ 
puting. R Foundation for Statistical Computing, Vienna, 
Austria. [Available from website, accessed January 
2018.] 
Rideout, R. M., and M. J. Morgan. 
2007. Major changes in fecundity and the effect on population 
egg production for three species of north-west Atlantic flat¬ 
fishes. J. Fish Biol. 70:1759-1779. 
Sampson, D. B., and S. M. Al-Jufaily. 
1999. Geographic variation in the maturity and growth 
schedules of English sole along the U.S. west coast. J. Fish 
Biol. 54:1-17. 
Sampson, D. B., and Y. W. Lee. 
1999. An assessment of the stocks of petrale sole off Wash¬ 
ington, Oregon, and Northern California in 1998, 99 p. 
Pac. Fish. Manage. Counc., Portland, OR. [Available from 
website.] 
Schneider, C. A., W. S. Rasband,, and K.W. Eliceiri. 
2012. NIH Image to ImageJ: 25 years of image analysis. Nat. 
Methods 9:671-675. 
Skjaeraasen, J. E., J. A. Devine, J. A. Godiksen, M. Fonn, H. OtterA, 
O. S. Kjesbu, B. Norberg, 0. Langangen, and 0. Karlsen. 
2016. Timecourse of oocyte development in saithe Pollachius 
virens. J. Fish Biol. 90:109-128. 
Starr, P. J., and J. Fargo. 
2004. Petrale sole stock assessment for 2003 and recommen¬ 
dations for management in 2004. Can. Sci. Advis. Seer., 
Res. Doc. 2004/036, 80 p. [Available from website.] 
Stawitz, C. C., T. E. Essington, T. A. Branch, M. A. Haltuch, 
A. B. Hollowed, and P. D. Spencer. 
2015. A state-space approach for detecting growth variation 
and application to North Pacific groundfish. Can. J. Fish. 
Aquat. Sci. 72:1316-1328. 
Stawitz, C. C., F. Hurtado-Ferro, P. Kuriyama, J. T. Trochta, 
K. F. Johnson, M. A. Haltuch, and O. S. Hamel. 
2016. Stock assessment update: status of the U.S. petrale sole 
resource in 2014, 210 p. Pac. Fish. Manage. Counc., Port¬ 
land, OR. [Available from website.] 
Turnock, J., M. Wilkins, M. Saelens, and C. Wood. 
1993. Status of West Coast petrale sole in 1993. In Status 
of the Pacific coast groundfish fishery through 1993 and 
recommended biological catches for 1994: stock assessment 
and fishery evaluation, p. G1-G42. Pac. Fish. Manage. 
Counc., Portland, OR. [Available from website.] 
Wallace, R. A., and K. Selman. 
1981. Cellular and dynamic aspects of oocyte growth in tele- 
osts. Am. Zool. 21:325-343. 
