June 9, 1923 
Rogue Types in Garden Peas 
831 
The increased variation of the Fj aa plants is probably due for the 
most part to crossing-over of the factors Yy for stipule shape and partly 
to the recombination of modifying factors affecting the same character. 
The mean ratio of the A segregates, 1.905^:0.0142, is significantly less 
than that of the Mummy parent, 2.088±0.0050, and the variability of 
the former is greater than that of the pure AA plants. The increased 
variability of the colored-flowering segregates, besides being accounted 
for largely by crossing-over, could, in addition, be due to a dissimilarity 
in stipule shape of AA and Aa plants. The heterozygous plants being 
in excess would tend to lower the mean of the population. Actually 
the Fi plants did appear to have stipules intermediate in shape between 
those of the two parents, but no measurements are available for compar¬ 
ison. 
It is realized that the number of Fg individuals measured is small but, 
taken alone, the statistical results would offer fairly good evidence of 
linkage between the factor or factors for stipule shape and the A factor 
for flower color. In connection with the Fg classification based on 
stipule shape and flower color, the fact of linkage between the Aa and 
Yy factors is well established. 
GRADUS rogue; X MUMMY 
The Gradus rogues used in the series of crosses with Mummy were all 
descended from a rogue plant. Si5, selected in 1916 from a commercial 
planting of Gradus. Progeny of the same rogue. Si5, when crossed with 
Gradus type behaved as typical rogues in inheritance (cultures 9.127a- 
9.1275, 9.1320, and 9.1321, Table II). 
The Fj plants (Table XII) of Gradus rogue X Mummy, and reciprocal 
had, with one exception, rogue stipules, and in this respect appeared 
similar to the pure rogues. As volunteer crossing occurred in the F^ of 
Gradus rogue x Gradus, it might be expected to take place among the 
Pj hybrids of rogue and Mummy. Since the Fj plants in the latter cross 
exhibited most of the dominant Mendelian factors present in any of the 
strains growing in the experimental gardens in 1919, it would be difficult 
to detect volunteer hybrids in the Fg generation (F^ out-crosses). How¬ 
ever, as the Fj hybrids were very late in flowering, cross-pollination by 
insects, if not entirely absent was cut down to a minimum. 
Tabi,^ Pedigrees of cultures of Gradus rogueXMumnty {P16) and reciprocal 
Culture. 
Pedigree. 
Mature plants. 
9.720 
(Si5-2-5r X P16-10)—I to 6. 
Rogues. 
9.721 
(Si5-2-i2r X P16-10)—I to 9. 
Do. 
9.722 
(Si5-2-3r X P16-10)—I to 7. 
Do. 
9-723 
(815-2-191X P16-10)—I to 6. 
Do. 
9-725 
(Si5-i-6r X P16-10)—I to 3. 
Do. 
9.726 
(Si5-i-9r X P16-10)—I to 8 . 
Do. 
9.727 
(Si5-3-ir X P16-10)—I to 8 . 
All rogues but No. 6, which 
resembled Fj of P16 x type. 
9.728 
(Si5-3-4r X P16-10)—I to 3. 
Rogues. 
9.729 
(Si5-3-5r X P16-10)—I to 8. 
Do. 
9-730 
(P16-12 X Si5-4-5r)—I to II. 
Do. 
Total. 69 
43324—23-3 
