July 1, 1924 
Inheritance in Kola-Hard Federation Grosses 
17 
Table XI.—-Segregation of 4,432 F 2 plants of the Kota-Hard Federation and 
reciprocal crosses into two classes for color of kernels, when grown at Davis } Calif., 
St. Paul, Minn., and Mandan, N. Dak., in 1922 
1 
j 
Num¬ 
ber of ! 
F, 
plants 
Number and percentage of F 2 plants 
Devia¬ 
tion 
from 
15:1 
Prob- 
Source and cross 
Red kernels 
White kernels 
Total 
number 
able 
error 
DAVIS, CALIF. 
KotaXHard Federation. 
11 
325 
Per cent 
92.9 
25 
Per cent 
7.1 
350 
3 
3. 05 
Hard FederationXKota.-. 
19 
643 
94.6 
37 
5.4 
680 
5 
4. 26 
Total_ 
30 
968 
94.0 
62 
6.0 
1,030 
2 
5.24 
ST. PAUL, MINN. 
KotaXHard Federation_ 
20 
615 
94.3 
37 
5.7 
652 
4 
4.17 
Hard FederationXKota_ 
40 
922 
96.1 
37 
3.9 
959 
23 
5.06 
Total_ 
60 
1,537 
95.4 
74 
4.6 
1,611 
27 
6. 52 
MANDAN, N. DAK. 
KotaXHard Federation_ 
12 
1,085 
94.9 
58 
5.1 
1,143 
13 
5.53 
Hard FederationXKota_ 
15 
626 
96.6 
22 
3.4 
648 
19 
4.16 
Total_ 
•27 
1,711 
95.5 
80 
4.5 
1,791 
32 
6.88 
ALL THREE STATIONS 
KotaXHard Federation_ 
23 
2,025 
94.4 
120 
5.6 
2,145 
14 
7. 59 
Hard FederationXKota_ 
45 
2,191 
95.8 
96 
4.2 
2, 287 
! 47 
7. 79 
Grand total_ 
68 
4,216 
95.1 
216 
4.9 
4,432 
1 61 
! 
10.86 
The data in Table XI show a fairly 
good agreement to a 2-factor or 15:1 
ratio. For this ratio 93.75 per cent 
should be red-kerneled and 6.25 per 
cent white. The grand total shows 
95.1 per cent red and 4.9 per cent white 
or a deviation from the 15:1 ratio of 
59 ±10.86, and, therefore, is not a 
close fit. A more detailed study of the 
data shows that in some cases there 
were no significant differences. 
There again is evidence of slight 
maternal influence on this character in 
the reciprocal crosses, as there is a 
consistant difference in the percentages 
at all three points. The numbers and 
percentages for the Kota X Hard Fed¬ 
eration cross show a close fit to the two- 
factor hypothesis at each of the three 
localities and in the total for all. The 
reciprocal cross, however, in wlpch 
Hard Federation is the female parent 
and which has the white or recessive 
type of kernel, shows a close fit at only 
one of the three localities, Davis, Calif., 
and not* in the total for all. 
A more certain separation of the 
kernels into the red and white classes 
was possible with the Davis, Calif., ma¬ 
terial than with that from St. Paul, 
Minn., and Mandan, N. Dak. Envi¬ 
ronmental conditions could account for 
the close fit which was obtained from 
the total material grown at Davis and 
for the significantly different results 
for the total material at St. Paul and 
Mandan. 
It would appear therefore that two 
Mendelian factors are involved in 
kernel color in this cross, and that the 
lack of a significantly close fit to the 
15:1 ratio in the reciprocal or Hard 
Federation X Kota cross and in the 
grand total is due to environmental 
conditions and possible slight maternal 
influence. When Kota, which has the 
dominant red kernels, was used as the 
female parent, a significantly close fit 
was obtained under all three different 
conditions. 
In order further to determine the 
factors involved, F 3 material of the 
1923 crop from Davis, Calif., was 
studied. The data obtained are given 
in Table XII. 
