July 1, 1924 
Inheritance in Kota-Hard Federation Crosses 
15 
With this character there is evidence 
of slight maternal influence on the 
reciprocal crosses as there is a consis- 
tant difference in the percentage at all 
three points. The percentages of 
white-glumed plants are the greater 
when Kota is used as the female parent 
and the percentages of brown-glumed 
plants are greater with Hard Federa¬ 
tion as the female parent. 
The totals of white-glumed plants 
in the reciprocal crosses at each of the 
three stations in the grand total show 
that there resulted a considerably 
larger number of plants having white 
glumes than would be expected on a 
3:1 basis in general, the ratio being 
nearer 2:1 than 3:1. A close fit to the 
expected 3:1 ratio was obtained for the 
Hard Federation X Kota cross at Davis 
and Mandan, where the deviations 
were 20 ±7.62 and 10 ±7.48, respec¬ 
tively. At St. Paul, the results from 
this cross also were much nearer a 
good fit than the reciprocal, although 
significantly different from the ex¬ 
pected ratio, which is true also for the 
total. It is important, however, that 
where Hard Federation, which has 
brown glumes and the dominant color, 
was used as the female parent, a close 
fit was obtained under two of the three 
different sets of conditions. 
It will be recalled that brown glumes 
in the Fi were a somewhat lighter 
brown than that of Hard Federation. 
Considerable variation was apparent 
in the F 2 regarding glumes classed as 
brown or white. The glumes of Kota 
are yellowish rather than white, but 
there appeared to be some white- 
glumed plants in the F 2 progeny. The 
differences, however, were not con¬ 
trasting enough to permit any more 
definite grouping, even with the ma¬ 
terial from Davis, Calif., where grow¬ 
ing conditions were ideal and the colors 
almost perfectly developed. At Man- 
dan and more particularly at St. Paul, 
summer rains and damage by rust 
made separations on the two colors 
difficult. The study was continued in 
the F 3 at Mandan and Davis and the 
data obtained are given in Table X. 
Table X. —-Segregation of 15,946 F 3 plants of the Kota-Hard Federation and recip¬ 
rocal crosses into two classes for color of glumes, when grown at Mandan, N. 
Dak., and Davis, Calif., in 1923 
F 2 families ; 
F 3 plants 
Source, F 2 classes, and segregation in the F 3 
Number 
1 
Percent* i 
age of 
class 
Percent¬ 
age of 
F 2 ° 
Glumes 
Brown 
. • 1 
White 
MANDAN, N. DAK. 
1 
Glumes white: 
White __ 
126 
81.8 
23.1 
, 2,918 
White and brown.....- 
28 
18.2 
5.2 
280 
360 
Total___ 
154 
100.0 
28.3 | 
280 
3,278 
Glumes brown: 
25.8 
18.5 
1,704 
Frown 
72 
Brown and white.. 
207 
74. 2 
53.2 
3,300 
1,657 
Total.... 
279 
100.0 
71. 7 
5,004 
1,657 
Mandan total.... 
433 
200.0 
100.0 
5,284 
4,935 
DAVIS, CALIF. 
Glumes white: 
89.3 
25.5 
2,118 
Whit.A _ 
75 
White and brown.... 
9 
10.7 
3.1 
! 156 
57 
Total_-.. 
84 
100.0 
28. 6 
156 
2,175 
Glumes brown: 
39.8 
28.4 
1,350 
Frown _ 
45 
Brown and white...-. 
68 
60.2 
43.0 
1,488 
558 
Total.— -.-. 
113. 
100.0 
71.4 
2,838 
558 
Davis total... 
197 
200.0 
100.0 
2,994 
2,733 
° Weighted in proportion to percentage of F 2 plants in each class of total for same station in 1922, given 
in Table IX. 
