Jan. 5, 1924 
Seed-Color Inheritance in Sorghum Crosses 
57 
and were classified with the glossy white phenotype with no nucellar 
layer. 
A number of selfed heads were obtained from the F 2 rows of Sunrise 
kafir X feterita and its reciprocal grown in 1921. Seed from these and 
from several unbagged heads were sown in head rows in 1922 to determine 
the behavior of the material in the F 3 generation. Descriptions of the 
seed from the F 2 heads and the phenotypic distribution of the F 3 progenies 
grown from them in 1922 are shown in Table II. 
Tabus II .—Description of F 2 seed heads of the cross, Sunrise kafir X feterita, and its 
reciprocal, and distribution of the progenies obtained from these heads into seed-color 
classes in IQ22 
' Distribution of F2 
progeny. 
Description of seed heads. 
j Brown nucellar 
! layer present. 
I 
Brown 
nucellar 
layer 
absent, 
white 
seeds. 
Factorial 
composition. 
Brown 
seeds. 
Chalky 
white 
seeds. 
Sunrise kafir X feterita (row 1 in 1921): 
1. Not bagged, white, no nucellar layer... 
0 
0 
40 
fbbss, 
{ bbSs, or 
2. Not bagged, brown, nucellar layer. 
86 
27 
0 
[bbSS 
BBsS 
3. Selfed, brown, brown nucellar layer... 
57 
25 
22 
BbSs 
4. Do. 
124 
O 
0 
BBSS 
5. Not bagged, white, brown nucellar layer 
0 
78 
23 
Bbss 
Sunrise kafir X feterita (row 2 in 1921): 
1. Selfed, brown, brown nucellar layer... 
74 
0 
27 
BbSS 
2. Do. 
43 
48 
BbSs 
3* Do. 
82 
30 
40 
BbSs 
4. Not bagged, white, no nucellar layer... 
0 
0 
123 
[bbss, 
{bbSs, or 
5. Selfed, brown, brown nucellar layer_ 
60 
23 
34 
IbbSS 
BbSs 
6. Do. 
114 
0 
33 
BbSS 
7. Selfed, white, brown nucellar layer_ 
0 
148 
0 
BBss 
Feterita X Sunrise kafir: 
1. Selfed, white, brown nucellar layer_ 
0 
90 
26 
Bbss 
2. Do. 
0 
79 
29 
Bbss 
3- Do...... 
0 
89 
0 
BBss 
4. Selfed, white, no nucellar layer. 
0 
0 
133 
[bbss, 
| bbSs, or 
5. Selfed, white, brown nucellar layer_ 
0 
94 
0 
(bbSS 
BBss 
6. Selfed, brown, brown nucellar layer_ 
85 
3 i 
0 
BBSs 
The factorial constitution of most of the F 2 heads used as seed can be 
determined definitely from the F 3 results. This is given in the last 
column of Table II, on the basis of the two-factor hypothesis previously 
outlined, and will account for all observed types. These two factors, 
therefore, not only account for the surface color of the kernel, but also 
for the presence or absence of the very apparent brown nucellar layer. 
The ratios obtained in the F 3 generation, while not exact, are as close 
to the expected as such small numbers of plants would require. Three 
possible factor combinations are suggested for the parent heads of the 
