58 
Journal of Agricultural Research 
Vol. XXVII, No. I 
three pure white F 3 progenies. Any of the factor combinations indicated 
for these F 2 heads would produce all white-seeded plants, without brown 
nucellar layer, in the F 3 generation, and consequently the specific one 
can not be determined. 
SUNRISE KAFIR X BUACKHUIX KAOUIANG 
A cross was made in 1919 between Sunrise kafir and Blackhull kaoliang, 
using Sunrise kafir as the pistillate parent. Reciprocal crosses were 
attempted, but the Blackhull kaoliang spikelets seemed to be very sus¬ 
ceptible to mutilation and the emasculated flowers dried without pro¬ 
ducing seed. Part of the crossed seeds were sown in 1920 and the 
remainder in 1921. 
The Fj kernels of the Sunrise kafir X Blackhull kaoliang cross were of 
a tan or brown surface color. A brown nucellar layer also was present. 
The similarity in appearance of feterita and Blackhull kaoliang kernels 
and the similarity in the color of the seeds of Fi crosses between these 
varieties and Sunrise kafir indicate that the same or very similar color 
factors are present in feterita and Blackhull kaoliang. As previously 
stated, the strain of kaoliang used probably is a selection from a natural 
hybrid between Barchet kaoliang and feterita. 
Seed from two selfed heads of the F t generation grown in 1920 were 
sown in head rows in 1921, and two similar head rows were grown in 
1922 from F t plants selfed in 1921. The data on seed color obtained 
from the F 2 progenies in 1921 and 1922 are presented in Table III. 
Table III. — Distribution of seed color in the F 2 generation of the cross Sunrise kafir X 
Blackhull kaoliang , grown in 1921 and 1922 
Row number. 
Y ear. 
1- 
f 
J Total 
| number. 
1 
Plants. 
With seeds havi 
Brown nucellar 
layer present. 
ng— 
Brown 
nucellar 
layer 
absent, 
seeds 
white. 
Seeds 
brown. 
Seeds 
white. 
I. 
1921 ! 
224 
132 
44 
48 
2 . 
I 9 21 
226 
132 
47 
47 
I. 
1922 
227 
133 
46 
48 
2. 
1922 
250 
159 
43 
48 
Total. 
927 
tr 
180 
17 A 
191 
i'll 
Calculated (0 : 2 : 4 ratio). 
jj" 
521 
7 . £ 
Deviation.. 
! 
/4 
6 
A T 
! 
j J 
4 A 
X 2 = 9.803. P—.0076, 1 in 131, or 0.76 per cent. 
The results indicate that the segregation of seed-color factors in the 
Sunrise kafir X Blackhull kaoliang cross is similar to that in the crosses 
between Sunrise kafir and feterita. While the data in Table III diverge 
somewhat widely from the calculated ratios, yet in view of similar data 
from other crosses the agreement is close enough to confirm the two- 
factor hypothesis. 
