Jan. 5, 1924 
Seed-Color Inheritance in Sorghum Crosses 61 
elusion seems justified that in the feteritaXRed kafir cross there are 
three factors influencing the color of seed, and that the three color factors 
are independently inherited, i. e., there is no evidence of linkage. 
The possible combinations of three factor pairs, Bb , S$, Rr , in the F 3 
generation are grouped under four phenotypes, as follows: 
Brown surface. 
White surface, 
Red or pink surface, 
r . ; 
i White surface, 
brown nucellar 
brown nucellar 
brown nucellar 
brown nucellar 
layer present. 
layer present. 
layer absent. 
layer absent. 
| 
i BBSSRR 
i BBssrr 
i bbSSRR 
| 1 bbSSrr 
2 BbSSRR 
2 Bbssrr 
2 bbSSRr 
2 bbSsrr 
2 BBSSRr 
! 3 
2 bbSsRR 
1 bbssrr 
4 BbSSRr 
4 bbSsRr 
! 4 
2 BBSsRR 
1 bbssRR 
I 
j 
4 BbSsRR 
i 
1 
2 bbssRr 
1 BBSSrr 
— 
2 BbSSrr 
4 BBSsRr 
8 BbSsRr 
2 BBSsrr 
4 BbSsrr 
1 BBssRR « 
2 BbssRR a 
2 BBssRr« 
12 
! 
! 
j 
• i 
j 
! 
i 
| 4 BbssRr a 
j 45 
a Under the climatic conditions occurring in 1922, there was no evidence of a phenotype similar to feterita 
but carrying red in the pericarp. It was concluded, therefore, that an R factor with a B factor in the presence 
of ss gave a seed which was classified with the phenotype having brown surface color and brown nucellar 
layer. However, in 1923, the F2 material from the same source as in 1922 showed a number of plants whose 
seeds possessed a brown nucellar layer and had a white outer color marked with red. These doubtless are 
individuals having the factorial constitution of BBssRR, BbssRR, BBssRr, or BbssRr. 
SUNRISE KAFIR X RED KAFIR 
In the discussion of the crosses previously described, in which feterita 
or Blackhull kaoliang was one parent and Sunrise or Red kafir the other 
parent, it was shown that brown seeds were obtained in the F x generation 
and in more than half of the variates in the F 2 generation. 
The cross Sunrise kafir X Red kafir was made in 1920. A few 
crossed seeds were sown in 1921, from which two selfed F t heads were 
obtained. The seed from these was sown in head rows in 1922, and 
the segregation for seed color in the F, generation of this cross is shown 
in Table VI. 
Table VI .—Distribution of seed-color in the F 2 generation of the cross Sunrise 
kafir X Red kafir , grown in IQ22 
Row number. 
I Total 
number of 
plants. 
Plants with 
i red and 
j tinted seed. 
Plants 
with white 
seed. 
1. 
144 
143 
! 
HI 
33 
42 
2. 
IOI 
Total. 
287 
212 
75 
72 
Expected (3:1 ratio). 
215 
Deviation, 34:4.95. p-g 1 - =0*606. 
