64 
Journal of Agricultural Research 
Vol. XXVIt, No. I 
In the F 2 generation of the feterita X Red kafir cross, four phenotypes 
are obtained, these being (a) brown with brown nucellar layer, (6) white 
with brown nucellus, (c) red and pink with no brown nucellus, and ( d ) 
white with no brown nucellus. The approximate ratio of these pheno¬ 
types is 45:3:12:4, indicating that three main factors influence the 
inheritance of seed color in this cross. 
In the crosses Sunrise kafir X Red kafir and White kafir X Red kafir, 
segregation in the F 2 generation occurs in a simple monohybrid ratio of 
three red-seeded plants to one white-seeded plant. 
Crosses between White kafir and Sunrise kafir show that the seed- 
color of these two varieties is genetically identical. 
The factors concerned in the inheritance of seed color in these sorghum 
crosses are B> a factor for brown nucellar layer, which produces brown 
color in the epidermis in the presence of S; b , its allelomorph, giving no 
brown nucellar layer when homozygous; 5 , a factor which will cause the 
development of color in the epidermis of the seed; s } its allelomorph, which, 
when present in the homozygous condition, will prevent the develop¬ 
ment of the brown color in the epidermis, R t a factor for dark red, which, 
when present, causes the development of red color in the epidermis; 
and r, its allelomorph, absence of red color. 
The factorial constitution of feterita and Blackhull kaoliang with 
respect to seed color is BBssrr , of Sunrise and White kafir, bbSSrr , 
and of Red kafir, bbSSRR. 
LITERATURE CITED 
(1) Conner, A. B., and Karper, R. E. 
1923. THE INHERITANCE OP SEED COAT COLOR IN CERTAIN CROSSES IN GRAIN 
sorghum. In Jour. Amer. Soc. Agron., v. 15, p. 338-344. Literature 
cited, p. 344. 
(2) Graham, R. J. Douglas. 
1916. POLLINATION AND CROSS-FERTILIZATION IN THE JUAR PLANT, (ANDRO- 
pogon sorghum, BROT.). In Mem. Dept. Agr. India Bot. Ser., v. 8, 
p. 201-216, 1 pi. 
(3) Harrington, George T., and Crocker, William. 
1923. STRUCTURE, PHYSICAL CHARACTERISTICS, AND COMPOSITION OF THE 
PERICARP AND INTEGUMENT OF JOHNSON GRASS SEED IN RELATION 
to its physiology. In Jour. Agr. Research, v. 23, p. 193-222, 10 fig. 
Literature cited, p. 221-222. 
(4) Harris, J. Arthur. 
1912. A SIMPLE TEST OF THE GOODNESS OF FIT OF MENDELIAN RATIOS. In 
Amer. Nat., v. 46, p. 741-745. 
(5) Harz, C. O. 
1885. landwirthshaftliche samenkunde. ix, 1362 p., 201 fig. Berlin. 
(6) Karper, R. E., and Conner, A. B. 
1919. natural cross-pollination in milo. In Jour. Amer. Soc. Agron., 
v. 11, p. 257-259. 
(7) Mitlacher, Wilhelm. 
1901. UEBER EINIGE EXOTISCHE GRAMINEENFROCHTE, DIE ZUR MENSCHLICHEN 
nahrung dienen. n. andropogoneae. In Ztschr. Allg. Oesterr. 
Apoth. Ver., Jahrg. 39, p. 831-836. 
(8) SiEglinger, John B. 
1921. CROSS-POLLINATION OF MILO IN ADJOINING ROWS. In JotlT. Amer. Soc. 
Agron., v. 13, p. 280-282. 
(9) Vinall, H. N., and Cron, A. B. 
1921. IMPROVEMENT OF SORGHUMS BY HYBRIDIZATION. In JoUT. Heredity, 
v. 12, p. 435 - 443 . 6 fig. 
(10) Winton, A. L. 
1903. THE ANATOMY OF THE FRUIT OF CERTAIN CULTIVATED SORGHUMS. In 
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