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Records of the Australian Museum (2017) Vol. 69 
Figure 11. Janthina janthina (Linnaeus), everted radula and odontophore in feeding position; enlarged copy of Graham (1965: fig. 5). 
(A) right lateral view; (B) dorsal view; (C) ventral view. Abbreviations: a, anterior transverse muscle; af accessory flexor muscle of the 
cartilage; bs, buccal septum; c, cartilage; dl, dorsal lip;/ anterior edge of foot;/ jaw; o, oesophagus; oa, opening of oesophagus on dorsal 
surface of extrovert; ol, opening of labial gland; op, outer posterior transverse muscle; ot, tip of odontophore; ps, protractor muscle of 
subradular membrane; rt, radular tooth; s, snout; sg, salivary gland; sm, subradular membrane; vl, ventral lip. Scale bar 10 mm. 
Colonna (1616; reprinted by Dollfus & Dautzenberg, 1932: 
pi. 13, lower fig.). The radulae of Recluzia species have not 
been described in detail, and are not visible protruding through 
the mouth in living specimens in the same way as Janthina 
radulae do, but presumably are identical to those of Janthina 
and benthic Epitoniidae. Feeding and prey are discussed 
further below. 
Epipodia. Photographs of the external anatomy of a living 
Janthina janthina animal in its normal life position, inverted 
beneath the float, were published in black-and-white by 
Wilson & Wilson (1956: pi. 1, fig. 1-2; Fig. 1) and again in 
colour by Yonge&Thompson (1976: pi. 5, lower). A drawing 
based on the first of these photographs was also published by 
Pinn (1980: fig. 5). Drawings showing similar details were 
published also by Okutani (1956: text-figs 1-10). These 
photographs, Okutani’s (1956) drawings, and further side 
views of the head of J. janthina published here (Figs 4D-E) 
clearly show the protruding snout (short and contractile, but 
not an eversible proboscis) and forked cephalic tentacles, the 
