Beu: Evolution of Janthina and Recluzia 
145 
Figure 14. Janthina fossil localities in the eastern Atlantic, showing location of Morocco, Madeira, the Canary Islands, Selvagem Grande 
Island, and the Azores Islands. • Janthina typica (possibly including other Janthina species at Madeira and Selvagem Grande I.). Localities: 
(1 ) Fouarat, Morocco (J. typica, type locality of Acrybia chouberti ); (2) La Esfinge, on La Isleta, Gran Canaria, Canary Islands (J. typica) -, 
(3) Selvagem Grande I. (J. typica or possibly J. chavani)-, (4) Sao Vicente, Madeira (J. typica or possibly J. chavani)-, (5) Santa Maria I., 
Azores (type locality of both Hartungia typica and Janthina krejcii sp. nov.). 
an intermediate volcano-sedimentary succession formed 
during 5.33^1.32 Ma (early Zanclean). The Touril Complex 
includes all the marine sedimentary rocks on the island 
other than late Pleistocene deposits. The Touril Complex is 
overlain in turn by a younger shield volcano, the Facho-Pico 
Alto Complex, formed during 4.32-3.94 Ma (late Zanclean), 
again without marine interbeds. Younger “parasitic” cones 
ceased erupting at only 2.84 Ma in the area of the Facho-Pico 
Alto Complex. This new dating indicates that all records of 
marine fossils from Santa Maria Island are early Zanclean 
in age (5.33^1.32 Ma), including the type localities of both 
Janthina typica and J. krejcii sp. nov. Figure 13 shows the 
new interpretation of Santa Maria Island stratigraphy by 
Sibrant et al. (2015). All Pliocene fossil localities (including 
those that have not yielded fossil Janthina specimens) 
and a few of the main late Pleistocene fossil localities are 
superimposed, but several of the newly recorded Pleistocene 
localities along the south coast are not shown. 
