Beu: Evolution of Janthina and Recluzia 
175 
Etymology. The new species is named in honour of Dr Karl 
Krejci-Graf, formerly of Universitat Frankfurt am Main, who 
collected the type material and advised the present writer 
about its locality. 
Time range. Zanclean, c. 4.8^1.3 Ma; the older limit is 
poorly constrained, but the upper limit on Santa Maria Island 
cannot be younger than 4.32 Ma (Sibrant et al., 2015). 
Janthina chavani (Ludbrook, 1978) 
Figs 27-28 
Heligmope postulatus (Bartrum).-Fleming, 1953b: 139 
(misidentification). 
Hartungia postulata (Bartrum).-Carter, 1972: 306, 321 
(misidentification). 
Hartungia typica typica (Bronn).- Johnstone et al., 1973: 14; 
Quilty, 1974a: 308; Quilty, 1974b: 29 (misidentification 
by A. Beu & G. Kendrick pers. comm.). 
Hartungia dennanti chavani Fudbrook, 1978: 119, pi. 12, 
figs 1-14; Fudbrook, 1983: 45, figs 3h-j; Fudbrook, 
1984: 232, figs 57o-p; Kendrick et al., 1991: 424, 436. 
Parajanthinajaponica Tomida & Itoigawa, 1982: 60, pi. 19, 
figs la-c; Ogasawara, 2002: 545 (in part). 
Hartungia japonica (Tomida & Itoigawa).-Tomida & 
Itoigawa, 1984: 112, pi. 31, figs la-2b; Tomida & 
Itoigawa, 1989: 126, pi. 23, figs la-2d;Noda etal., 1995: 
83, figs 11.7a-d; Nobuhara et al., 1995: 39, figs 3.2a-b; 
Tomida & Kitao, 2002: 158, figs 2.1a-2c; Ogasawara, 
2002: 394, 545. 
Hartungia chavani (Fudbrook).-Kendrick in Tomida & 
Itoigawa, 1984: 112, pi. 31, figs 3a-5b; Beu & Maxwell, 
1990: 411; Maxwell in Spencer etal, 2009: 245. 
Hartungia sp.-Nobuhara et al, 1995: 38, figs 3.1a-b. 
Kaneconcha knorri Kaim, Tucholke & Waren, 2012: 247, 
figs 3A-E. 
Janthina (Hartungia ) typica (Bronn).-Tomida et al, 2013: 
60, figs 3E-F only (in part misidentified). 
Type material. Hartungia dennanti chavani, holotype 
WAM69.300c, with six figured and numerous unfigured 
paratypes in WAM, Geological Survey of Western Australia, 
and Geological Survey of South Australia (listed by 
Ludbrook, 1978: 120; WAM and GSSA material observed); 
from Roe Calcarenite (late Pliocene; Beu & Darragh, 
2001: 31, fig. 6), pits c. 50-125 km W of Eucla, Roe Plain, 
southeastern Western Australia. As noted above, although 
Beu & Darragh (2001: fig. 6) suggested an early Piacenzian 
age, correlation with successions in New Zealand now 
indicates a late Piacenzian age for Roe Calcarenite. Ludbrook 
(1978: 120) also recorded specimens from water wells 
beneath Perth, Western Australia, from Plio-Pleistocene 
Bridgewater Limestone in the region around Naracoorte, 
South Australia, and from Reedy Wells, Culburra, South 
Australia (the last not seen; from Bridgewater Limestone 
near Mount Gambier). 
Parajanthina japonica, holotype MFM110004, from 
Dainichi Sand (late Pliocene, Piacenzian, upper part of 
planktonic foraminiferal zone N21), Higashigumi, Iida, 
Shizuoka Prefecture, Honshu, Japan (Tomida & Itoigawa, 
1982: 61); not seen. 
Kaneconcha knorri, holotype (Figs 28A, D, G) in Institute 
of Paleobiology, Polish Academy of Sciences, Warsaw, ZPAL 
Ga.16/1, with six incomplete paratypes, ZPAL Ga. 16/2-7, 
from Knorr dredge station 180-2-26,23°23'N 45°23'W, Kane 
Megamullion, east flank of Adam Dome, mid-Atlantic ridge, 
3293-2827 m (mapped by Kaim et al., 2012: fig. 1). The 
writer has seen photographs of only the holotype, which is 
a slightly crushed specimen of Janthina chavani. Although 
it has weak spiral folds on the weakly convex sutural ramp, 
it is identified as J. chavani partly because the matrix was 
dated by calcareous nannofossils by M.-P. Aubry (the leading 
expert on nannofossil biostratigraphy; in Kaim et al., 2012) 
as zone NN16B, 2.5-2.8 Ma (late Piacenzian-earliest 
Gelasian, latest Pliocene-earliest Pleistocene). Kaim et al. 
(2012: 427^129, figs 5C-D) mentioned that although six 
other specimens were collected, they are all fragmentary. 
They described two shell layers (both now replaced by 
calcite) in a paratype fragment, and suggested that the outer, 
dark brown later 25 pm thick is a periostracum preserved by 
replacement with calcite. The inner layer, 175 pm thick over 
the columella, is milky white homogeneous calcite. However, 
reidentification of this shell as J. chavani demonstrates that 
the outer layer is the brown calcite outer shell layer (violet 
in life), only 25 pm thick, whereas the inner white layer is 
the original aragonite inner layer. In most specimens from 
other localities the inner layer apparently is not so thick. 
Other material examined. Australia: South Australia: Two paralecto- 
types of Heligmope dennanti, from Hallett Cove Sandstone, coast east of 
Hallett Cove, south of Adelaide (see above under H. dennanti', SAMA 
T1515 A, B); Geological Survey of South Australia, Bridgewater Limestone 
(Piacenzian-Calabrian?), Kanawinka Fault scarp, Naracoorte, SE South 
Australia (GSSA F87/65,2; photographs sent by N. H. Ludbrook); cliffs S 
of car park. Point Ellen, Vivonne, Kangaroo Island, South Australia, grid ref. 
Vivonne 970138 (locality PL3173, NMVP316448,22); thin cemented beds 
in Bridgewater Limestone (late Piacenzian-Calabrian?) at top of Henske’s 
Quarry, Elderslie Road, 2.6 km SE ofNaracoorte, SE South Australia, grid 
reference Hynam 793085 (locality PL3249, NMV P318105, 20, in small 
limestone blocks, collected by T. A. Darragh and A. G. Beu; also block of 
specimens c. 1 m 2 observed at quarry office); Naracoorte quarries, SE of 
Naracoorte, South Australia (clearly from Henske’s Quarry; NMV P316446, 
1). Western Australia: Roe Plain: Roe Calcarenite (late Piacenzian), 
district around Hampton microwave repeater tower. Roe Plain, 48-126 
km W of Eucla Motel, SE Western Australia, suites in several museums 
(WAM.71-1438a-g, 7; WAM.69-298, 1; WAM.69-300a-z, 26; WAM.69- 
301a-d, 4; WAM.69-299a-d, 4; WAM.59-305, 1; WAM.69-302a-c, 3; 
WAM.69-303,1; WAM.70-2156a-c, 3; WAM.69-297a-f, 6; WAM.67-778a, 
b, 2; WAM.69-306, 1; WAM.69-304, 1; NMV P26917, 2; P316447, 18; 
P322322,1, Fig. 271; GNS WM14468,10, Figs 27A-H, 28C, F, J); Madura 
Cave, Roe Plain (WAM.62-50, 1; WAM.63.44, 1). Perth Basin: “lower” 
Ascot Formation, water wells in Perth Basin, collected over many years by 
G. W. Kendrick (Kendrick in Quilty, 1974b: 29; Kendrick etal, 1991: 424, 
436), all in WAM; most lots consist of one specimen or fragment: 30 m. 
Vale’s bore, Evelyn St, Gosnell’s, Perth (WAM.70-2615); Redcliffe primary 
school bore, Perth (WAM.69-292, WAM.69-293, WAM.69-294); Geo¬ 
logical Survey of Western Australia bore Gnangara no. 21, W Bullsbrook, 
Perth (WAM.68-179, many fragments); Kowalski’s bore, corner Bullfinch 
& Balfour Streets, Gosnell’s, Perth (WAM.69-296). 
New Zealand: Mangapanian (late Piacenzian-earliest Gelasian): Hawke’s 
Bay: Cricklewood Road (GS12515, W19/f020, grid ref. W19/798357; 4 
fragments); brown sandstone 3 km upstream from road bridge, Mohaka 
River (GS13079, W19/f031, grid ref. W19/634313; 1, now fragmentary); 
Matahorua Road, Tutira (GS12508, V19/f011, grid ref. V19/481224; 1); 
sandstone between conglomerate beds, Pohokura Road, Tutira (GS12507, 
V20/f018, grid ref. V20/444168; 2 fragments). Whanganui Basin: basal 
conglomerate of Komako Formation, Te Ekaou Stream, Pohangina Valley 
(OUGD, OU8037, T23/f6565, grid ref. T23/576173; 1; Carter, 1972). 
Nukumaruan (Gelasian-earliest Calabrian): Hawke’s Bay: Darkies Spur 
Formation, road cut. Darkies Spur Road, Arapaoanui Valley, c. 30 km N 
of Napier (GS11225, V20/f8002, grid ref. V20/407104; 1 + fragment; 
Figs 281, K). Whanganui Basin: Komako Formation, Pohangina Valley 
(Carter, 1972: 306, 321), Makawakawa Stream (OUGD, OU7597, T23/ 
f6516, grid ref. T23/594099; 1); Konewa Stream (OUGD, OU7668, T23/ 
f6548, grid ref. T23/595201; 1); Te Ekaou Stream (OUGD, OU8125, T23/ 
f6563, grid ref. T23/574175; 1); Hautawa Shellbed, Te Ekaou Stream, Dept, 
of Earth Sciences, University of Waikato (T23/f060, grid ref. T23/577174; 
