178 
Records of the Australian Museum (2017) Vol. 69 
1); lowest lens of Nukumaru Limestone, Waitotara “desert”, coast W of 
Whanganui (GS4258, R22/f6488, grid ref. R22/581489; 1; Fleming, 1953b: 
139); 0.1 m-thick sandy shell lens in massive mudstone, “undifferentiated 
Upper Okiwa Group” between Tuha Sand and Ohingaiti Sand (Fleming, 
1953b: 133, 136), road cut on hillside 200 m E of Makohine Stream, 2 km 
S of Ohingaiti, Rangitikei Valley (T22/f8506, grid ref. T22/550356; GNS 
TM4495,1, Figs 28B, E, H; D. Cowe collection, 1967, several); Tewkesbury 
Formation (late Nukumaruan), shellbed enclosing Vinegar Hill Tephra (MIS 
61,1.75Ma;Pillans etal., 2005: 79,figs5A, 11;Townsend etal., 2008:fig. 
35), Brunswick Road, SE side Kai Iwi Valley, W of Whanganui (GS15348, 
R22/f6542A, grid ref. R22/773506; 1). 
The only other material observed in world museums is the 
type material of Hartungia dennanti chavani, listed above. 
Distribution. Janthina chavani is particularly abundant 
in the type area, in Roe Calcarenite (late Piacenzian) on 
the Roe Plain, southeastern Western Australia. It is also 
abundant at a few quarries in Bridgewater Limestone (late 
Piacenzian-Calabrian?) around Naracoorte, South Australia, 
and Tate’s two paralectotypes of Heligmope dennanti from 
the upper part of Hallett Cove Sandstone near Hallett Cove 
in South Australia are also J. chavani. Ludbrook (1978) also 
recorded it from a few other localities in South Australia. 
Ludbrook (1983,1984) also recorded./ chavani from Point 
Ellen Formation at Cape Jervis, Fleurieu Peninsula, mainland 
South Australia, and at Point Ellen, Vivonne, Kangaroo 
Island, South Australia. Ludbrook (1983; 45, figs 3h-j; 
Geological Survey of South Australia GSSA10025a-c, three 
illustrated) recorded 17 specimens from Point Ellen and 
four from Cape Jervis; further specimens have since been 
collected by T. A. Darragh at Point Ellen (listed above). Many 
fragments and a few complete specimens have also been seen 
from “lower” Ascot Formation in water wells in the Perth 
Basin, Western Australia (material inWAM). In New Zealand 
it is much less common and widespread than J. typica, 
and is recorded from only 13 localities in Mangapanian 
and Nukumaruan (late Piacenzian-early Calabrian) rocks 
in Hawke’s Bay and Whanganui Basin. In Japan, a few 
specimens have been collected from near the Pacific coast 
of SE Honshu Island, and two were reported by Tomida et 
al. (2013) from Hioki, Miyazaki Prefecture, near the east 
coast of Kyushu. The single Atlantic record is from Kane 
Megamullion, on the mid-Atlantic ridge (Kaim etal., 2012), 
where specimens apparently were dredged from “normal” 
seabed, and not from a hydrothermal seep site as Kaim et 
al. (2012) thought. Late Pliocene-early Pleistocene rocks of 
suitable facies for the preservation of Janthina apparently are 
not exposed on the Atlantic islands where J. typica occurs, 
although the records from Sao Vicente, Madeira, and from 
Selvagem Grande Island require re-collection to be certain 
of their identities and are possibly J. chavani. 
Dimensions. See Table 5. 
Diagnosis. Teleoconch moderate-sized to very large for 
Janthina (up to H 48, D 40 mm), covered with fine, straight, 
closely spaced axial ridges; axial ridges tending to fade out 
over last whorl of large specimens; 8-11 spiral folds per 
whorl (9 or 10 on most specimens) but most specimens 
with spiral folds significantly less obvious than on J. typica, 
at least two spiral folds suppressed on upper sutural ramp. 
Outer lip sinus small, basal, as in J. typica, but slightly 
wider, narrowly V-shaped in some specimens. Teleoconch 
increasing in height with weak allometry; most juvenile 
specimens with low spires, most large specimens taller and 
narrower than all other large Janthina species, although much 
shorter and wider than Recluzia species. 
Table 5. Dimensions of Janthina chavani ^dimensions 
from original publications; specimens with very weak spiral 
folds indicated within parentheses; NSF, number of spiral 
folds). Stage abbreviations: Wm, Mangapanian New Zealand 
Stage (late Piacenzian-early Gelasian, 3.0-2.4 Ma); Wn, 
Nukumaruan New Zealand Stage (Gelasian-early Calabrian, 
2.4-1.63 Ma). 
locality 
height 
diam. 
NSF 
H/D 
Hartungia dennanti chavani holotype* 
44 
38 
— 
1.16 
Kaneconcha knorri holotype* 
32 
34.5 
— 
0.93 
Parajanthina japonica holotype* 
31.2 
37.3 
11 
0.84 
Australian specimens, all from Roe Plain: 
88 km W of Eucla Motel (WAM) 
33.1 
29.6 
9 
1.12 
88 km W of Eucla Motel 
27.0 
25.9 
9 
1.04 
88 km W of Eucla Motel 
36.0 
29.3 
9 
1.23 
88 km W of Eucla Motel 
47.9 
37.9 
9 
1.26 
88 km W of Eucla Motel 
44.0 
37.4 
(8) 
1.18 
88 km W of Eucla Motel 
44.8 
35.6 
8 
1.26 
88 km W of Eucla Motel 
40.8 
35.4 
8 
1.15 
88 km W of Eucla Motel 
32.1 
27.3 
9 
1.18 
88 km W of Eucla Motel 
29.2 
25.7 
10 
1.14 
88 km W of Eucla Motel 
30.1 
27.1 
10 
1.11 
88 km W of Eucla Motel 
38.3 
33.0 
9 
1.16 
88 km W of Eucla Motel 
37.8 
35.5 
9 
1.06 
88 km W of Eucla Motel 
36.4 
31.5 
9 
1.16 
88 km W of Eucla Motel 
26.7 
27.0 
9 
0.99 
88 km W of Eucla Motel 
33.0 
29.6 
11 
1.11 
88 km W of Eucla Motel 
32.4 
33.2 
9 
0.98 
88 km W of Eucla Motel 
38.1 
38.1 
(8) 
1.0 
88 km W of Eucla Motel 
30.8 
30.4 
9 
1.01 
88 km W of Eucla Motel 
24.6 
23.6 
11 
1.04 
88 km W of Eucla Motel 
41.0 
37.5 
9 
1.09 
88 km W of Eucla Motel 
40.7 
39.1 
9 
1.04 
126 km W of Eucla Motel (WAM) 
44.8 
37.8 
(8) 
1.19 
126 km W of Eucla Motel 
28.7 
28.6 
9 
1.00 
126 km W of Eucla Motel 
29.4 
26.6 
9 
1.11 
126 km W of Eucla Motel 
42.8 
41.5 
(10) 
1.03 
126 km W of Eucla Motel 
30.5 
28.0 
9 
1.09 
84 km W of Eucla telegraph station (WAM) 40.3 
32.4 
8 
1.24 
84 km W of Eucla 
35.8 
34.9 
9 
1.03 
84 km W of Eucla 
37.3 
33.5 
9 
1.11 
84 km W of Eucla 
35.2 
31.5 
9 
1.12 
84 km W of Eucla 
29.6 
28.6 
8 
1.03 
74 km W of Eucla Motel (WAM) 
24.3 
26.8 
9 
0.91 
74 km W of Eucla Motel 
35.8 
31.5 
10 
1.14 
68 km W of Eucla Motel (WAM) 
23.3 
24.7 
10 
0.94 
48 km W of Eucla Motel (WAM) 
35.1 
32.2 
9 
1.09 
48 km W of Eucla Motel 
21.6 
22.2 
9 
0.97 
48 km W of Eucla Motel 
31.4 
28.3 
10 
1.11 
26 km W of Madura (WAM) 
28.7 
28.7 
(9) 
1.0 
26 km W of Madura 
26.8 
28.5 
9 
0.94 
26 km W of Madura 
37.9 
34.6 
10 
1.10 
Hampton microwave tower (WAM) 
22.8 
23.7 
10 
0.96 
640 m N of Hampton microwave (WAM) 
25.7 
28.3 
9 
0.91 
NMV P322322, 88 km W of Eucla Motel 
37.7 
33.3 
(9) 
1.13 
GNS WM14468, 16 km W of Madura 
47.8 
41.3 
(6) 
1.16 
GNS WM14468, 16 km W of Madura 
42.4 
39.3 
(8) 
1.08 
GNS WM14468, 16 km W of Madura 
42.1 
33.9 
(7) 
1.24 
GNS WM14468, 16 km W of Madura 
37.4 
33.4 
(10) 
1.12 
GNS WM14468, 16 km W of Madura 
35.1 
33.2 
(10) 
1.06 
GNS WM14468, 16 km W of Madura 
33.85 
31.3 
(11) 
1.08 
GNS WM14468, 16 km W of Madura 
32.6 
29.5 
(9) 
1.11 
GNS WM14468, 16 km W of Madura 
27.5 
27.8 
(5) 
0.99 
GNS WM14468, 16 km W of Madura 
27.6 
27.3 
(8) 
1.01 
New Zealand specimens: 
T23/f6565, Te Ekaou Stream, OUGD, Wm 
24.5 
30.4 
10 
0.81 
T22/f8506, above Makohine viaduct, Wn 
23.9 
28.0 
(8) 
0.85 
T22/f8506, above Makohine Viaduct 
20.6 
26.5 
9 
0.78 
T22/f8506, above Makohine Viaduct 
23.8 
25.7 
9 
0.93 
GS4258, Nukumaru Limestone, Wn 
27.0 
27.4 
(5) 
0.99 
OU8037, T23/f6563, Te Ekaou Stream, Wn 13.3 
16.2 
9 
0.82 
OU8125, T23/f6563, Te Ekaou Stream, Wn 20.5 
25.3 
9 
0.81 
