Bean, Revision of Anisomeles 
331 
Fig. 6. Diagrams of infloresence types in Anisomeles. A. entirely monochasial. B. once dichasial at base. C. twice 
dichasial at base. 
cyme results in a lax slender inflorescence 
where each flower or fruit is clearly visible. 
The ‘twice-dichasiaf cyme results in a 
compact sub-globose inflorescence where the 
branching pattern is difficult to see and many 
of the flowers or fruits are obscured. The 
‘once-dichasiaf cyme is of an intermediate 
form. Inflorescence characters have been 
assessed using fruiting material, where the 
internodes are fully expanded, and it is easier 
to detect the patterns involved. The number 
of flowers per monochasium is typically 
quite variable within species, but is of some 
diagnostic value. The distance between the 
individual flowers or fruits (it is the fruiting 
distance that is measured in this paper) is 
diagnostic for some species. Adjacent fruits 
can be as much as 12 mm apart, giving rise to 
a very lax cyme, or as little as 0.7 mm apart, 
resulting in a congested cyme. 
5. Corolla colour, shape and indumentum 
(Figs. 1A, ID, 7A-C). 
The corolla is predominantly pink or mauve 
to purple for most species with areas of white 
usually present. In Anisomeles heyneana , the 
corolla is often pure white. The descriptions of 
flower colour given on herbarium labels are so 
varied or generalised that it makes any analysis 
of the character quite pointless. The corolla 
shape is remarkably consistent throughout 
the genus; it consists of a small entire upper 
lip, adjacent to or in contact with the stamens; 
and a large lower lip, generally somewhat 
recurved. The two terminal lobes are obtuse 
and mostly fused, with an emarginate apex 
indicating their fusion; the lateral lobes are 
similarly obtuse and scarcely extend past the 
base of the lateral lobes. Between the lateral 
lobes is an area here dubbed the ‘platform’, 
a roughly rectangular area bounded by the 
lateral lobes, terminal lobes, and the throat. 
The platform is sometimes flat, but often 
somewhat raised or puckered, and in some 
species, it bears relatively long stiff hairs. The 
number of these hairs is of diagnostic value. 
Within the corolla tube is the annulus, which 
is surmounted by many tiny stiff erect hairs. 
6. Fruiting calyx morphology 
The calyx in Anisomeles grows in size from 
the budding stage to anthesis and through 
to fruiting, and measurements of the calyx 
near anthesis will vary greatly depending on 
the exact stage of floral development. All of 
the features of the flowering calyx are also 
present on the fruiting calyx, and are more 
easily seen and assessed on the latter, and 
measurements are more reliable as the calyx 
has by then stopped growing. 
The fruiting calyx offers a number of very 
diagnostic morphological characters, which 
have been assessed and measured from dried 
pressed material at the fruiting stage, where 
the nutlets (Fig. 7H) are mature or nearly so. 
