Bean, Revision of Anisomeles 
343 
13 Stems with patent hispid hairs; plants procumbent; external hairs on 
fruiting calyx 1.1-2.4 mm long. 14. A. languida 
13. Stems without patent hispid hairs; plants erect; external hairs on fruiting 
calyx 0.6-1 mm long. 25. A. vulpina 
14 Nutlets 1.5-1.7 mm long; cauline leaves with 19-33 pairs of marginal lobes 21. A. papuana 
14. Nutlets 1.8-2.2 mm long; cauline leaves with 3-18 pairs of marginal lobes. 15 
15 Corolla platform glabrous; petioles 21-38% of lamina length. 19. A. moschata 
15. Corolla platform with 20-100 hairs; petioles 13-22% of lamina length . . 26. A. xerophila 
I. Anisomeles ajugacea (F.M.Bailey & 
F.Muell.) A.R.Bean comb, nov.; Teucrium 
ajugaceum F.M.Bailey & F.Muell., Synop. 
Queensl. FI. Second Suppl. 48 (1888). Type: 
Australia: Queensland. Cook District: 
Musgrave Electric Telegraph Station, 
undated, T. Bard ay-Millar s.n. (holo: BRI 
[AQ340432], 2 sheets). 
Anisomeles sp. (Big Coleman River 
J. R.Clarkson+ 7119); Henderson (2002). 
Procumbent shrub, 0.05-0.2 m high. Upper 
stems and rachises with patent hispid hairs; 
short curved hairs absent or retrorse, sparse; 
stalked glandular hairs absent; sessile glands 
8-32 mm 2 . Cauline leaves 16-29 mm long, 
14-24 mm wide, 1.1-1.4 times longer than 
wide, base narrowly cuneate (< 60°) or 
attenuate; marginal lobes dentate, irregular 
or regular, 2-4 on each side, acute, 2.5-5.1 
mm deep; petioles 0-3 mm long, 0-13% 
of lamina length. Lamina upper surface 
indumentum appressed, eglandular, 1-2.2 
mm long, moderately dense (hairs 0.1-0.2 
mm apart) or dense (hairs < 0.1 mm apart), 
sessile glands 8-112 mm 2 ; lower surface 
indumentum appressed, eglandular, 1-2.5 
mm long, moderately dense (hairs 0.1-0.2 mm 
apart) or dense (hairs < 0.1 mm apart), sessile 
glands 8-96 mm 2 ; transition from leaves to 
floral bracts gradual. Floral bracts ovate or 
broadly ovate, 5-13 mm long, 3-7 mm wide, 
not consistently exceeding verticils. Verticils 
(inflorescence clusters) all separated on 
rachis, cymes entirely monochasial, with 3-5 
flowers per monochasium, peduncles 0-3 mm 
long on lowermost cluster; bracteoles obovate 
or spathulate, 5-10 mm long, 1-3 mm wide. 
Corolla tube same length as calyx; annulus 
3.3-5 mm from base of corolla, annulus 
hairs 0.4-0.5 mm long; upper lip elliptical, 
4-6.6 mm long, with glandular hairs on outer 
surface or with eglandular hairs on outer 
surface; lower lip pink, 5.6-67 mm long to 
end of lateral lobes, 9.8-11.3 mm long overall, 
glabrous on platform. Longest stamens 12-14 
mm long from base of corolla tube; filament 
hairs 0.35-0.5 mm long, mainly along middle 
part or mainly at distal end. Style 12.7-15.5 
mm long; longer stigma lobe 0.85-1.2 mm 
long, shorter stigma lobe 0.4-0.65 mm long. 
Fruiting calyces 2-5 mm apart on rachilla; 
fruiting calyx narrowly campanulate, 7-8 mm 
long, 3.4-5 mm wide at lobe apices, 1.5-2.1 
times longer than wide, exterior surface with 
all hairs same size and type, hairs eglandular, 
1.5-2.1 mm long, sessile glands 48-128 mm 2 ; 
lobes acute, 2.6-3 mm long. Fruiting calyx 
fringe hairs about the same length throughout, 
0.2-0.35 mm long at apical end, 0.1-0.25 mm 
long at sinus end, sinus hairs absent, inner 
surface of tube glabrous. Nutlets 2.3-2.6 mm 
long. Figs. 3A, 8A. 
Additional selected specimens examined : Australia: 
Queensland. Cook District: Near Little Laura River, 
SSW of Laura, Jul 1990, Bean 1906 (BRI); 28.3 km from 
the Kennedy River on the Jedda Creek track to King 
River Station, Jun 1981, Clarkson 3717 (BRI, K, MO, 
PERTH); 2 km S of the Big Coleman River on the Coen 
to Musgrave Road, May 1987, Clarkson 7119 & Simon 
(BRI, NSW); 3.5 km N of South Five Mile Creek on the 
Peninsula Developmental Road, Apr 1991, Clarkson 
8929 & Neldner (BRI, K); 27 km N of Musgrave, Cape 
York Peninsula, Jun 2004, Foley s.n. (BRI [AQ610577]); 
21.4 km E of Musgrave, May 2004, Gray 8910 (BRI, 
CANB, CNS, NSW); Musgrave Electric Telegraph 
Station, May 1891, Jacobson s.n. (BRI [AQ161262]); 
25 km by road towards Jowalbinna from Peninsula 
Development Road, Jul 2007, McDonald KRM6826 & 
Johnson (BRI); 6.7 km along Pormpuraaw Road from 
Gulf Development Road junction near Musgrave, May 
2010, McDonald KRM9188 (BRI, CNS); N of Musgrave 
on Cape York Development Road, just S of Red Blanket 
Creek, May 2004, Wannan BSW3496 & Graham (BRI); 
