Bean, Revision of Anisomeles 
347 
0.25-0.3 mm long, moderately dense (hairs 
0.1-0.2 mm apart), sessile glands 8-48 mm 2 ; 
lower surface indumentum of erect glandular 
hairs 0.25-0.3 mm long, moderately dense 
(hairs 0.1-0.2 mm apart), sessile glands 32- 
80 mm 2 ; transition from leaves to floral bracts 
abrupt. Floral bracts elliptical or ovate, 5-15 
mm long, 2-4.5 mm wide, not consistently 
exceeding verticils. Verticils (inflorescence 
clusters) overlapping near apex, cymes twice 
dichasial (± globose), with 4-7 flowers per 
monochasium, peduncles 0-1 mm long on 
lowermost cluster; bracteoles spathulate or 
linear, 6.4-11 mm long, 0.8-1.6 mm wide. 
Corolla tube shorter than calyx; annulus 3.4-4 
mm from base of corolla, annulus hairs 0.25- 
0.35 mm long; upper lip ovate or elliptical, 
4.5-4.8 mm long, with glandular hairs on 
outer surface; lower lip 7-7.2 mm long to end 
of lateral lobes, 12-13.4 mm long overall, with 
more than 100 eglandular hairs on platform. 
Longest stamens 12-13 mm long from base 
of corolla tube; filament hairs 1.2-1.3 mm 
long, mainly at distal end. Style 12.5-13.5 
mm long; longer stigma lobe 0.65-0.75 mm 
long, shorter stigma lobe 0.55-0.7 mm long. 
Fruiting calyces 0.7-1.1 mm apart on rachilla; 
fruiting calyx cylindrical, 11.4-12.6 mm 
long, 3.2-4 mm wide at lobe apices, 3.1-3.6 
times longer than wide, exterior surface with 
all hairs same size and type, hairs glandular, 
0.25-0.35 mm long, sessile glands 8-32 mm 2 ; 
lobes acute, 3.3-4.4 mm long. Fruiting calyx 
fringe hairs about the same length throughout, 
0.15-0.25 mm long at apical end, 0.15-0.25 
mm long at sinus end, sinus hairs absent, 
inner surface of tube with sparse long hairs 
in medial section or glabrous. Nutlets 2.3-2.4 
mm long. Figs. 3C, 8D. 
Additional specimens examined: Australia: Northern 
Territory. Mt Bundey, Arnhem Highway, Mar 1987, 
Dunlop 6884 & Wightman (DNA). 
Distribution and habitat: Anisomeles 
bundeyensis is endemic to the Northern 
Territory. It is known only from Mt Bundey 
and nearby Mt Bundey East, SE of Darwin 
(Map 2). It grows on granite outcrops. 
Phenology : Flowers are recorded for March 
and May; fruits recorded in May. 
Notes: Anisomeles bundeyensis is 
distinguished by the abundant glandular hairs 
on the stems, the very long bracteoles, the 
twice-dichasial cymes on the verticils, and 
the long fruiting calyces. It is closely related 
to A. viscidula, but differs by the lack of 
retrorse eglandular hairs and hispid hairs on 
the stems, the considerably larger corolla, the 
twice-dischasially branched cymes and the 
fruiting calyces only 1-1.5 mm apart. 
Conservation status: This species has a very 
restricted Area of Occupancy (<20 km 2 ), and 
is known from just two locations. A status of 
Vulnerable, criterion D2, is recommended. 
Etymology: Named for the location of Mount 
Bundey. 
5. Anisomeles candicans Benth. in Wall., 
PI. Asiat. Rar. 1: 59 (1830); Epimeredi 
candicans (Benth.) Rothm., Repert. Spec. 
Nov. Regni Veg. 53: 12 (1944). Type citation: 
Yenangheun, ripae Irawaddi. Type: Burma. 
Yenanghuen [Yenangyaung], ripae Irawaddi, 
in 1826, [Wallich Cat. No. 2038] (lecto [here 
designated]: K 000846316, image!). 
Anisomelesmalabaricavm. nigrescens Benth. 
in Wall., PI. Asiat. Rar. 1: 59 (1830). Type 
citation: ad ripas Irawaddi. Types: Burma. 
Prome, in 1826, [Wallich Cat. No. 2037] (syn: 
K); Irawaddi River, [1826], [Wallich Cat. No. 
2037/4] (syn: K; syn: BM 000984304). 
Shrub, height unknown. Upper stems and 
rachises without patent hispid hairs; short 
curved hairs retrorse, moderately dense; 
stalked glandular hairs absent; sessile glands 
8-32 mm 2 . Cauline leaves 65-91 mm long, 15- 
32 mm wide, 2.6-4.3 times longer than wide, 
base narrowly cuneate (< 60°) or attenuate; 
marginal lobes crenate or dentate, irregular or 
regular, 11-18 on each side, acute or obtuse, 
0.6-2.1 mm deep; petioles 9-17 mm long, 13- 
20% of lamina length. Lamina upper surface 
indumentum erect or curved, eglandular, 
0.3-0.4 mm long, moderately dense (hairs 
0.1-0.2 mm apart), sessile glands 48-80 mm 2 ; 
lower surface indumentum erect or curved, 
eglandular, 0.4-0.6 mm long, dense (hairs 
< 0.1 mm apart), sessile glands 8-32 mm 2 ; 
transition from leaves to floral bracts abrupt. 
Floral bracts lanceolate or elliptical, 6-45 
