354 
bracteoles spathulate or linear, 4.3-9.5 mm 
long, 0.5-1.3 mm wide. Corolla tube longer 
than calyx or the same length as calyx; 
annulus 3.1-4 mm from base of corolla, 
annulus hairs 0.2-0.3 mm long; upper lip 
elliptical, 37-5.9 mm long, with eglandular 
hairs on outer surface; lower lip 4.8-76 mm 
long to end of lateral lobes, 8.5-14.5 mm 
long overall, platform glabrous or with 1-20 
eglandular hairs. Longest stamens 12.5-14.5 
mm long from base of corolla tube; filament 
hairs 0.5-1.4 mm long, mainly at distal end. 
Style 13-15 mm long; longer stigma lobe 0.4- 
0.7 mm long, shorter stigma lobe 0.3-0.55 
mm long. Fruiting calyces 1-1.5 mm apart 
on rachilla; fruiting calyx cylindrical, 8.7-11 
mm long, 2.8-4.3 mm wide at lobe apices, 
2.3-37 times longer than wide, exterior 
surface with all hairs same size and type, 
hairs eglandular, longest ones 0.25-0.6 mm 
long, sessile glands 8-112 mm 2 ; lobes acute, 
1.8-3.3 mm long. Fruiting calyx fringe hairs 
about the same length throughout, 0.15-0.35 
mm long at apical end, 0.15-0.4 mm long at 
sinus end, sinus hairs absent, inner surface of 
tube with dense ring of long hairs in medial 
section or with sparse long hairs in medial 
section or glabrous. Nutlets 1.8-2.1 mm long. 
Figs. 3G, 4A, 8H, 9C 
Additional specimens examined : Australia: Northern 
Territory. Koolpin Gorge, Mar 1990, Brennan Brel90 
(DNA); 20 km SE of Twin Falls, May 1980, Craven 
5880 (CANB); Mt Brockman, Feb 1973, Dunlop 3323 
(BRI, CANB, DNA, NT); SE of Mt Howship, Arnhem 
land, Feb 1984, Dunlop 6628 & Russell-Smith (DNA); 
Nourlangie Rock, along path to lookout, Feb 1987, 
Gartrell & Cunliffe UNSW19974 (BRI, CANB, DNA); 
14.5 km WNW of Twin Falls, May 1980, Lazarides 
9106 (DNA, CANB, DNA); walking trail, Obiri Rocks, 
Apr 1987, Purdie 3175 (CANB); top of Twin Falls, 
Kakadu NP, Jun 1983, Russell-Smith 711 (DNA); 5 km 
E of Winwuyurr Creek crossing. Kakadu NP, Feb 1984, 
Russell-Smith 1102 (DNA); 6 km S of Oenpelli, Jul 
1983, Russell-Smith 1167 (DNA); 10 km SW of Oenpelli 
Aboriginal Settlement, May 1988, Weher 9890 (AD, 
BRI, DNA). 
Distribution and habitat : Anisomeles 
grandibractea is endemic to the Northern 
Territory. It is confined to the eastern parts 
of Kakadu NP, and the Oenpelli area of 
Arnhem Land (Map 7). It inhabits sandstone 
plateaux and gorges, in eucalypt woodland or 
Acacia scrub, and is often recorded from the 
Austrobaileya 9(3): 321-381 (2015) 
margins of monsoon vine forest in sandstone 
gorges, with at least some sites dominated by 
Allosyncarpia ternata S.T.Blake. Soils are 
skeletal or sandy. 
Phenology : Flowers are recorded between 
February and July; fruits from March to July. 
Notes : This species is characterised by 
seemingly axillary inflorescences, due to the 
floral bracts greatly exceeding the verticils. 
One collection (Lazarides 9106 ) has antrorse 
stem hairs, while other specimens have 
retrorse hairs on the stem. 
Anisomeles grandibractea has two forms. 
The typical form has relatively shorter and 
broad floral bracts, often brown tomentum 
on new growth, long hairs (0.45-0.7 mm 
long) on the upper leaf surface, long petioles 
(27-40% of lamina length), and obtuse or 
broadly cuneate leaf bases. The other form 
(represented by e.g. Lazarides 9106, Dunlop 
6628 & Russell-Smith ) has very long floral 
bracts, no brownish tomentum, short hairs 
(0.2-0.35 mm long) on the upper leaf surface, 
shorter petioles (16-28% of lamina length), 
and narrowly cuneate leaf bases. Further 
study may reveal these forms to be separate 
species. 
Conservation status : Least Concern. 
Etymology : The epithet alludes to the floral 
bracts of this species, which consistently 
exceed the verticils. 
11. Anisomeles heyneana Benth. in 
N.Wallich, PI. Asiat. Rar. 1: 59 (1830); 
Anisomeles secunda O.Kuntze, Revis. Gen. PI. 
2: 512 (1891), nom. illeg:, Epimeredi secundus 
Rothm., Repert. Spec. Nov. Regni Veg. 53: 
12 (1944), nom. illeg. Type: India, undated, 
B. Heyne s.n. [Wallich Cat. No. 2028] (lecto 
[here designated]: K 001057386, image!). 
Teucrium secundum Heyne, In Numer. List 
[Wallich] n. 2028 (1829), nom. nud. 
Erect or spreading shrub, 1.2-3 m high. 
Upper stems and rachises without patent 
hispid hairs; short curved hairs retrorse, 
sparse; stalked glandular hairs absent; sessile 
glands 8-16 mm 2 Cauline leaves 72-116 mm 
long, 30-38 mm wide, 2.4-3.1 times longer 
