Bean, Revision of Anisomeles 
355 
than wide, base narrowly cuneate (< 60°) or 
attenuate; marginal lobes dentate or serrate, 
irregular or regular, 12-22 on each side, 
acute, 0.5-2.3 mm deep; petioles 17-43 mm 
long, 24-37% of lamina length. Lamina upper 
surface indumentum appressed, eglandular, 
0.6-0.7 mm long, sparse (hairs > 0.2 mm 
apart), sessile glands 8-48 mm 2 ; lower surface 
indumentum erect or curved, eglandular, 
0.6-0.8 mm long, confined to veins or sparse 
(hairs > 0.2 mm apart), sessile glands 48-128 
mm 2 ; transition from leaves to floral bracts 
abrupt. Floral bracts ovate, 3.5-14 mm long, 
1.5- 7.5 mm wide, not consistently exceeding 
verticils. Verticils (inflorescence clusters) all 
separated on rachis, cymes once dichasial at 
base then monochasial, with 4-8 flowers per 
monochasium, peduncles 8-39 mm long on 
lowermost cluster; bracteoles spathulate or 
linear, 2.5-4.8 mm long, 0.3-1.2 mm wide. 
Corolla tube same length as calyx; annulus 
2.5- 3.5 mm from base of corolla, annulus 
hairs 0.1-0.15 mm long; upper lip elliptical, 
4.1-4.9 mm long, with eglandular hairs on 
outer surface; lower lip 5-5.5 mm long to 
end of lateral lobes, 9-10 mm long overall, 
glabrous on platform or with 1-20 eglandular 
hairs on platform. Longest stamens 11-12 mm 
long from base of corolla tube; filament hairs 
0.7-1.1 mm long, mainly along middle part. 
Style 12-13 mm long; longer stigma lobe 0.6- 
0.8 mm long, shorter stigma lobe 0.15-0.5 
mm long. Fruiting calyces 1.3-3 mm apart on 
rachilla; fruiting calyx cylindrical, 8-9.1 mm 
long, 37-4.5 mm wide at lobe apices, 1.9- 
2.2 times longer than wide, exterior surface 
with hairs of different sizes or types, hairs 
glandular and hairs eglandular, 0.4-0.9 mm 
long, sessile glands 48-112 mm 2 ; lobes acute, 
2.5- 3 mm long. Fruiting calyx fringe hairs 
about the same length throughout or longer at 
sinus end than at apical end, 0.2-0.3 mm long 
at apical end, 0.2-0.8 mm long at sinus end, 
sinus hairs absent or present, inner surface of 
tube with sparse long hairs in medial section. 
Nutlets 1.8-2.1 mm long. Figs. 4B, 9D. 
Western Hill catmint. 
Additional specimens examined : India. Karjat, North 
Konkan, Jan 1949, Fernandes 26 (A); near Dudh 
Sagor, Nov 1949, Fernandes 5430 (K); Londa, Bombay 
Presidency, Jan 1950, Fernandes 1012 (K); Nil-Gherries, 
1857-58, Perrottet (G); Hosangadi, Karnataka State, Jan 
1939, Raja 6340 (MH); below Phonda Ghat, Jan 1853, 
Ritchie 540 (E); near Kanheri Caves [between Thane & 
Borivali], Bombay presidency, Oct 1945, Sinclair 4562 
(E); Concan [Konkan], undated. Stocks s.n. (BRI, G, L, 
MH, NY, P). 
Distribution and habitat : Anisomeles 
heyneana is endemic to India. It occurs close 
to the west coast (Western Ghats), from 
Hosangadi to Mumbai (Map 6). None of 
the specimen labels include information on 
habitat; however, Efloraofindia (2007-2015) 
gives its habitat as “near forest clearing on 
hills and slopes”. 
Phenology : Flowers are recorded for January, 
October, and November; fruits in January. 
Notes : Bentham, in the protologue, 
distinguished Anisomeles heyneana 
by the secund (one-sided) pedunculate 
inflorescences, the long ascending branches 
and the small leaves. It is true that in A. 
heyneana , one of the cymes at each verticil 
sometimes does not develop forming a 
secund inflorescence, but this is not a reliable 
difference. 
Anisomeles heyneana is most likely to be 
confused with the sympatric A. indica , from 
which it differs by the narrower and often 
smaller leaves, the verticils all separated 
along the rachis, the long peduncle of the 
lower verticils, the white corolla, and the 
glabrous or sparsely hairy corolla platform. 
Conservation status: Data Deficient. 
12. Anisomeles indica (L.) Kuntze, Revis. 
Gen. PI. 2: 512 (1891); Nepeta indica L., 
Sp. PI. 2: 571 (1753); Epimeredi indicus (L.) 
Rothm., Repert. Spec. Nov. Regni Veg. 53: 12 
(1944). Type: Habitat in India (lecto: LINN 
726.28, fide Cramer 1981). 
Ballota disticha L., Mant. PI. 1: 83 (1767); 
Ajuga disticha (L.) Roxb., Hort. Bengal. 44 
(1814); Anisomeles disticha (L.) B.Heyne ex 
Roth, Nov. PI. Sp:. 254 (1821); Nepeta disticha 
(L.) Blume, Bijdr. 823 (1826). Type: Habitat 
in India (lecto: LINN 1211, fide Cramer 1981). 
