Bean, Revision of Anisomeles 
361 
patent hispid hairs; short curved hairs 
retrorse, moderately dense or dense; stalked 
glandular hairs absent; sessile glands 8-80 
mm 2 Cauline leaves 26-49 mm long, 12-25 
mm wide, 2-3.3 times longer than wide, 
base narrowly cuneate (< 60°) or attenuate; 
marginal lobes dentate or serrate, irregular or 
regular, 4-10 on each side, acute or obtuse, 
0.9-1.5 mm deep; petioles 4-13 mm long, 11- 
33% of lamina length. Lamina upper surface 
indumentum erect or curved, eglandular, 
0.4-1 mm long, sparse (hairs < 0.2 mm apart) 
or moderately dense (hairs 0.1-0.2 mm apart), 
sessile glands 8-64 mm 2 ; lower surface 
indumentum erect or curved, eglandular, 
0.4-07 mm long, confined to veins or 
moderately dense (hairs 0.1-0.2 mm apart), 
sessile glands 64-112 mm 2 ; transition from 
leaves to floral bracts gradual. Floral bracts 
16-46 mm long, 8-17 mm wide, consistently 
exceeding verticils. Verticils (inflorescence 
clusters) all separated on rachis, cymes once 
dichasial at base then monochasial, with 3-7 
flowers per monochasium, peduncles 1-6 mm 
long on lowermost cluster; bracteoles obovate 
or spathulate, 4-13 mm long, 0.7-5 mm wide. 
Corolla tube longer than calyx, or same 
length as calyx; annulus 2.2-3 mm from base 
of corolla, annulus hairs 0.1-0.3 mm long; 
upper lip ovate or elliptical, 4.1-6.2 mm long, 
with glandular hairs on outer surface; lower 
lip 6.2-77 mm long to end of lateral lobes, 
14-15.5 mm long overall, glabrous or with 
1-20 eglandular hairs on platform. Longest 
stamens 13-15 mm long from base of corolla 
tube; filament hairs 0.9-1.3 mm long, mainly 
at distal end. Style 13-15 mm long; longer 
stigma lobe 0.65-0.8 mm long, shorter stigma 
lobe 0.45-0.5 mm long. Fruiting calyces 
1-2 mm apart on rachilla; fruiting calyx 
cylindrical, 7-10 mm long, 4.3-5.5 mm wide 
at lobe apices, 17-2.5 times longer than wide, 
exterior surface with hairs of different sizes 
or types or with all hairs same size and type, 
hairs eglandular, 0.4-1.9 mm long, sessile 
glands 32-96 mm 2 ; lobes acute, 2-3 mm 
long. Fruiting calyx fringe hairs about the 
same length throughout, 0.15-0.3 mm long at 
apical end, 0.15-0.25 mm long at sinus end, 
sinus hairs absent, inner surface of tube with 
dense ring of long hairs in medial section or 
with sparse long hairs in medial section or 
glabrous. Nutlets 2.1-2.5 mm long. Figs. 4F, 
7E, 9H. 
Additional selected specimens examined: Australia: 
Queensland. Cook District: Alma-den, undated, Bick 
176 (BRI); Springmount Station, c. 13 km from the 
Mareeba - Dimbulah Road, on the road to Collins Weir, 
Apr 1983, Clarkson 4664 (BRI, CNS, DNA, K, MEL, 
MO, PERTH); 10 km from Alma-den on road to Mt 
Surprise, Jan 1992, Forster PIF9593 (BRI); 16 km from 
Mt Garnet on road to Lappa, 1 km past Mt Cardwell 
turnoff, Jan 1993, Forster PIF12777 & Bean (BRI, CNS, 
MEL); Pannikin Springs area, 29 km W of Mungana, 
Jan 1993, Forster P1F13013 & Bean (BRI, DNA); 
Copperfield River, Kidston Goldmine water supply dam, 
Gilbert Range, Feb 1994, Forster PIF14902 & Bean 
(BRI, CNS, MEL); Donkey Spring Creek, BulleringaNP, 
80 km NW of Mt Surprise, Apr 1998, Forster PIF22414 
& Booth (BRI); Stannary Hills, 11 km S of Mutchilba, 
Portion 603, May 2006, Forster PIF31593 & McDonald 
(BRI); Chillagoe, Jan 1931, Hubbard 6815 & Winders 
(BRI); Kent Holdings, W of road from Mt Garnet to 
Minnamoolka, May 1999, Jago 5239 & Wannan (BRI); 
1 km from Walsh River crossing, N of Chillagoe, Mar 
2000, McDonald KRM329 (BRI); near Barwidgi Road 
turnoff, S of Mt Garnet, Apr 2004, McDonald KRM2176 
& Covacevich (BRI); 51 km along Alma-den road from 
junction with Gulf Development Road, near Mt Surprise, 
May 2004, McDonald KRM2596 (BRI); Undara NP, 5.1 
km along northern boundary firebreak with St Ronan’s 
station, Dec 2006, McDonald KRM6032 (BRI); Gorge 
Creek, Mareeba, Apr 1962, McKee 9257 (CANB); 
Pinnacle Springs Road, 4 km W from Kennedy Highway, 
Apr 2004, Purdie 5924 (BRI, CANB); 2 km downstream 
of junction of Graves and Fulford Creeks, Burlington 
Station, N of Mt Surprise, Jun 1999, Thompson SLT2467 
& Newton (BRI); 100 mile swamps, ‘Rosella Plains’, Jul 
1981, Williams 81134 (BRI). North Kennedy District: 
12 km along Deadmans Road, off Silver Valley Road, 
Feb 1996, Forster P1F18415 & Ryan (BRI, CNS, NSW); 
c. 1.2 km ESE of Hidden Valley township, Paluma Road, 
Feb 2000, Pollock ABP853 & Edginton (BRI). 
Distribution and habitat : Anisomeles lappa 
is endemic to Queensland. It is found in the 
north-east of the state, including the western 
parts of the Atherton Tableland, Bulleringa 
NP, Alma-den, and Undara NP (Map 2). 
It grows in eucalypt woodland. Almost all 
records are from granite hills or sandy soils 
adjacent to granite outcrops. 
Phenology : Flowers and fruits are recorded 
from December to July inclusive. 
Notes: Anisomeles lappa can be distinguished 
by the prostrate or procumbent habit, the 
abundant patent hispid hairs on flowering 
stems, the short cauline leaves, and the long 
