Bean, Revision of Anisomeles 
363 
2013, Westaway JOW4142 (BRI, CANB, DNA); Bynoe 
Harbour, c. 200 m from MacKenzie Arm boat ramp off 
Barramundi Drive, Mar 2006, Wirf247 (DNA). 
Distribution and habitat: Anisomeles 
leucotricha is endemic to the Northern 
Territory, from Melville Island to Kakadu 
NP and Litchfield NP (Map 3). It grows 
on a variety of habitats, including vine 
thicket on foreshore, open woodland with 
Corymbia foelscheana (F.Muell.) K.D.Hill 
& L. A. S. Johnson and/or Erythrophleum 
chlorostachys (F.Muell.) Baill. on hills. 
Substrate is quaternary sand or sandstone. 
Phenology : Flowers are recorded from 
January to July; fruits between April and July. 
Notes: Anisomeles leucotricha is 
distinguished by the elliptical leaves with an 
attenuate base and a short even indumentum, 
the exceptionally long calyx fringe hairs, the 
closely arranged fruiting calyces (0.8-1.2 
mm apart), the adjacent verticils sometimes 
overlapping to form a dense cluster, and 
the tiny floral bracts. The leaves have many 
crenate lobes. The lower leaf surfaces have 
short antrorse hairs, very often conspicuously 
white, especially along the veins. 
This species possibly intergrades with 
A. brevipilosa to the east of (and west of) 
Katherine, as some specimens from these 
areas are difficult to allocate to either species. 
Conservation status: Least Concern (IUCN 
2012 ). 
Etymology: The epithet is from the Greek 
leucos - white, and trichos - hair or trichome. 
In this species, the hairs on the underside of 
the leaf are frequently conspicuously white, 
especially along the veins. 
17. Anisomeles macdonaldii A.R.Bean sp. 
nov. abundantia pilorum glandularium et 
absentia pilorum eglandularium retrorsorum 
caulibus insidentibus, foliis basalibus latis 
basi obtusa et petiolo longo (longitudine 32- 
54% laminae aequante) praeditis distinguitur. 
Typus: Australia: Queensland. Cook District: 
c. 8 km by road E of Chillagoe, 21 April 2013, 
K.R. McDonald KRM14083 & G.P. Guymer 
(holo: BRI; iso: MEL, distribuendi). 
Anisomeles salviifolia var. hispida Domin, 
Biblioth. Bot. 89: 567 (1928). Type: Australia: 
Queensland. Cook District: Smelling Bluff 
near Chillagoe, February 1910, K. Domin 
s.n. (syn: PR 530815); limestone hill near 
Chillagoe, February 1910, K. Domin s.n. (syn: 
PR 530816). 
Erect or spreading shrub, 0.4-2 m high. 
Upper stems and rachises with patent hispid 
hairs; short curved hairs absent or retrorse, 
sparse to moderately dense; stalked glandular 
hairs abundant; glandular hairs extending 
to lower stems; sessile glands 8-80 mm 2 
Cauline leaves 51-91 mm long, 32-63 mm 
wide, 1.2-1.9 times longer than wide, base 
obtuse or broadly cuneate (> 60°); marginal 
lobes crenate or dentate, regular, 10-18 on 
each side, acute or obtuse, 0.9-2.4 mm deep; 
petioles 19-40 mm long, 32-54% of lamina 
length. Lamina upper surface indumentum 
of erect glandular and curved eglandular 
hairs, 0.15-1.3 mm long, moderately dense 
(hairs 0.1-0.2 mm apart) or dense (hairs < 
0.1 mm apart), sessile glands 8-48 mm 2 ; 
lower surface indumentum of erect glandular 
and curved eglandular hairs, 0.15-1.3 mm 
long, moderately dense (hairs 0.1-0.2 mm 
apart) or dense (hairs < 0.1 mm apart), sessile 
glands 8-80 mm 2 ; transition from leaves to 
floral bracts gradual. Floral bracts ovate or 
broadly ovate, 14-47 mm long, 11-28 mm 
wide, not consistently exceeding verticils or 
consistently exceeding verticils. Verticils 
(inflorescence clusters) all separated on 
rachis, cymes entirely monochasial or once 
dichasial at base then monochasial, with 4-11 
flowers per monochasium, peduncles 2-28 
mm long on lowermost cluster; bracteoles 
spathulate or linear, 2.5-5.5 mm long, 0.4-1.1 
mm wide. Corolla tube longer than calyx, or 
same length as calyx; annulus 3-3.4 mm from 
base of corolla, annulus hairs 0.2-0.3 mm 
long; upper lip ovate or elliptical, 5.8-67 mm 
long, with glandular hairs on outer surface 
or with eglandular hairs on outer surface; 
lower lip pink, 8.2-9 mm long to end of 
lateral lobes, 14-16.5 mm long overall, with 
1-20 or 20-100 eglandular hairs on platform. 
Longest stamens 12.8-13.5 mm long from 
base of corolla tube; filament hairs 1.3-1.6 
mm long, mainly at distal end. Style 12.8-14 
