Bean, Revision of Anisomeles 
365 
dense (hairs < 0.1 mm apart), sessile glands 
8-48 mm 2 ; lower surface indumentum lanate, 
tangled or erect or curved, eglandular, 0.9-1.2 
mm long, very dense, obscuring surface at 
x40 magnification; transition from leaves to 
floral bracts abrupt. Floral bracts lanceolate 
or elliptical, 6-17 mm long, 2-5 mm wide, 
not consistently exceeding verticils. Verticils 
(inflorescence clusters) all overlapping, 
forming continuous terminal inflorescence or 
overlapping near apex, cymes once dichasial 
at base then monochasial or twice dichasial (± 
globose), with 5-14 flowers per monochasium, 
peduncles 0-1 mm long on lowermost cluster; 
bracteoles linear, 4.2-6.5 mm long, 0.3-0.5 
mm wide. Corolla tube shorter than calyx; 
annulus 3.5-4.2 mm from base of corolla, 
annulus hairs 0.1-0.15 mm long; upper lip 
elliptical, 4.5-5.2 mm long, with eglandular 
hairs on outer surface; lower lip 5-5.4 mm 
long to end of lateral lobes, 8.5-10.7 mm 
long overall, with 20-100 eglandular hairs 
on platform. Longest stamens 13-14 mm 
long from base of corolla tube; filament hairs 
0.4-0.6 mm long, mainly along middle part. 
Style 13-13.5 mm long; longer stigma lobe 
0.7-1 mm long, shorter stigma lobe 0.3-0.5 
mm long. Fruiting calyces 1.5-2.2 mm apart 
on rachilla; fruiting calyx cylindrical, 8-12.2 
mm long, 2.8-37 mm wide at lobe apices, 
3-4.3 times longer than wide, exterior surface 
with hairs of different sizes or types, hairs 
glandular and eglandular, 1.2-1.5 mm long, 
sessile glands 8-16 mm 2 ; lobes attenuate, 
3.1-5.3 mm long. Fruiting calyx fringe hairs 
about the same length throughout or longer 
at sinus end than at apical end, 0.3-0.6 mm 
long at apical end, 0.5-0.7 mm long at sinus 
end, sinus hairs present, inner surface of 
tube with dense ring of long hairs in medial 
section. Nutlets 1.9-2.2 mm long. Figs. 41, 
10C. Malabar catmint. 
Additional specimens examined : India. San Thome, in 
1856, Cleghorn s.n. (E); Kulasegaram, S. Travancore, 
Feb 1934, Erlanson 5378 (NY); Pondicherry, May 
1837, Gaudichand s.n. (P); Cuddalore, Nov 1959, 
Govindarajalu 3699 (L); Bounds of Vedanthangal 
Sanctuary, Tamil Nadu, Jan 1976, Henry 47027 
(MH); Tiruchi, Nov 1978, Matthew et al. RHT19392 
(L); Banavar - Arsikere road, Jan 1970, Saldanha 
16079 (E); Kanjamalai, Apr 1944, Sinclair 3464 (E); 
Papanasam project area, Madras State, Jun 1964, 
Subramanian 1653 (L); Perur, Noyal riverbank, Jun 
1956, Subramanyam 45 (MH); way to Pachakumatchi, 
Jun 1959, Subramanyam 8259 (L); Susindram, Tamil 
Nadu, Dec 1980, Swaminathan 68985 (MH); Walayar 
Dam, outlet canal, Kerala State, May 1964, Vajravelu 
19064 (MH); Peninsula Ind. orientalis, undated, Wight 
2173 (NY); Pulicat, Madras, Mar 1837, Wight 2173 (E). 
Sri Lanka. Kaddaikadu, Mar 1973, Bernardi 14265 (L); 
beside Trincomalee - Kinniya Road, Sep 1974, Cramer 
4325 (E); Palaiyutta, Mar 1994, Cramer 6960 et al. 
(E); 9 miles [14 km] NNE of Jaffa, along road to Palai, 
Dec 1974, Davidse 9090 & Sumithraarachchi (K, L); 
Trincomalee, Oct 1976, Fosberg 56390 (E, NY, P); Foul 
Point, Trincomalee district, Feb 1972, Jayasuriya 651 et 
al. (E); Talankuda, Batticaloa district, Apr 1973, Stone 
11188 (L); Just N of Trincomalee, Mar 1973, Townsend 
73/245 (E); Kinniyai, Trincomalee district, Sep 1974, 
Waas & Tirvengadum 809 (L, NY). Malaysia. Penang, 
in 1822, Wallich 2037/2 (NY). Western Indian Ocean. 
Mauritius, undated, Sieber s.n. (NY). 
Distribution and habitat : Anisomeles 
malabarica is native to Sri Lanka and 
southern India (Map 6). In addition, it is (or 
was) naturalised on Penang Island, Malaysia, 
and on the island of Mauritius. It inhabits 
open sunny areas; sandy flats on dunes near 
the coast, stream banks, and “waste ground”. 
Phenology : Flowers are recorded for every 
month of the year; fruits are recorded in 
January, April, September and December. 
Nomenclature : The authorship fox Anisomeles 
malabarica has often been given as “(L.) R.Br. 
ex Sims”, but the notation ‘Brown mss’ after 
the description in the protologue indicates 
that the author was Brown and not Sims. 
Hence the correct authorship is “(L.) R.Br. in 
Sims” or, where the plant name stands alone, 
just “(L.) R.Br”. 
Notes: Anisomeles malabarica is a very 
distinctive species because of its very densely 
tomentose stems, relatively narrow leaves, 
long attenuate calyx lobes and long hairs on 
the external surface of the calyx. 
Baker (1877) accepted A. malabarica as 
being native on Mauritius, but it seems far 
more likely that it was transported there by 
man, as specimens of the Mauritian plant are 
identical to those from India, and the species 
has no means of long-range seed dispersal. 
