Bean, Revision of Anisomeles 
369 
calyces 1-1.6 mm apart on rachilla; fruiting 
calyx cylindrical, 6.2-8.3 mm long, 3.1-3.5 
mm wide at lobe apices, 2.1-2.5 times longer 
than wide, exterior surface with all hairs same 
size and type, hairs eglandular, 0.2-0.6 mm 
long, sessile glands 8-160 mm 2 ; lobes acute, 
1.4-2.7 mm long. Fruiting calyx fringe hairs 
longer at sinus end than at apical end, 0.1- 
0.25 mm long at apical end, 0.45-1.1 mm long 
at sinus end, sinus hairs absent, inner surface 
of tube with dense ring of long hairs in medial 
section or glabrous. Nutlets 1.5-1.7 mm long. 
Figs. 5D, 10F. 
Additional specimens examined: Indonesia. Moluccas: 
Yamdena Island, road from Saumlakki to Olilit, Mar 
1938, Buwalda 4043 (L). Papua New Guinea. Central 
Province: Baroka, Mekeo district, Apr 1933, Brass 3715 
(BRI); Roger’s airstrip, c. 8 miles [13 km] W of Kanosia 
Plantation, Jul 1962, Darbyshire 653 (BRI, CANB, 
L); Nebiri Quarry, Port Moresby sub-district, Apr 
1970, Gebo 369 (CANB); near Jackson’s Airport, Port 
Moresby, Aug 1963, Heyligers 1006 (CANB); Tovobada 
Hills, 12 miles [19 km] N of Port Moresby, May 1965, 
Heyligers 1180 (BRI, CANB, L); S coast near Kwikila, 
Abau subdistrict, Jun 1969, Paijmans 767 (CANB); 
Rubulogo Creek, c. 18 miles [29 km] N of Port Moresby, 
Apr 1967, Pullen 6689 (CANB); north vicinity of Rigo, 
Aug 1962, Schodde 2700 (BRI, CANB); Port Moresby, 
Jul 1918, White 61 (BRI). Gulf Province: Near Malalaua, 
Mar 1966, Craven & Schodde 904 (CANB); Malalaua, 
lower Tauri River area, Feb 1966, Pullen 6528 (CANB). 
Morobe Province: Bulolo, Feb 1950, Fryar 3948 (BRI, 
CANB); Bulolo, Wau subdistrict, Jul 1970, Streimann 
& Kairo NGF47869 (BRI, CANB). Northern Province: 
Between Dabora and Wabubu, Cape Vogel Peninsula, 
Apr 1953, Brass 21876 (CANB). Western Highlands 
Province: Baiyer River, Nov 1954, Floyd & Womersley 
6826 (BRI). Western Province: Near Rouku, Morehead 
subdistrict, Jul 1973, Henty NGF49713 (BRI, CANB, 
L). Australia: Queensland. Cook District: Thursday 
Island, Jun 1897, Bailey s.n. (BRI [AQ160154]); Darnley 
Island, Torres Strait, Apr 2005, Hacks LAH285 (BRI, 
CANB); Yorke Island, Torres Strait, Oct 1971, Lawrie 
s.n. (BRI [AQ3927]); Mabuyag Island, Torres Strait, Aug 
2008, McKenna SGM222 (BRI, CANB); Gabba island, 
Torres Strait, Feb 2000, Waterhouse BMW5655 (BRI, 
CANB). 
Distribution and habitat: Anisomeles 
papuana is widely distributed in lowland New 
Guinea, also Torres Strait, Queensland, and 
the southern Moluccas Islands of Indonesia 
(Map 5). It inhabits savannah woodland at 
altitudes from 0-100 metres and mostly below 
30m (except the three specimens mentioned 
below). 
Phenology: Flowers and fruits are recorded 
from February to November inclusive. 
Notes: Anisomeles papuana is distinguished 
by the many lobes per leaf, the relatively short 
petioles, the consistently short cylindrical 
calyces and the nutlets only 1.5-1.7 mm long. 
Three specimens {Floyd & Womersley 
6826, Fryar 3948 , Streimann & Kairo 
NGF47869 ) are known from relatively high 
altitude (from 750 to 1150 metres) in Papua 
New Guinea. These specimens have a more 
compact infructescence, and the floral bracts 
exceed the verticils; they may represent an 
additional undescribed taxon. 
Conservation status : Least Concern. 
Etymology : The epithet refers to Papua 
New Guinea, where the great majority of 
collections have been made. 
22. Anisomeles principis A.R.Bean sp. 
nov. pilis antrorsis retrorsisve plusminusve 
patentibusve, eglandularibus, densis in 
caulibus, bracteolis latis, pilis 1-20 labio 
inferiori corollae insidentibus et pilis 
marginalibus brevibus loborum calycis 
distinguitur. Typus: Australia: Western 
Australia. South-west Osborne Island, 
Bonaparte Archipelago, 29 June 1973, P.G. 
Wilson 11152 (holo: PERTH). 
Erect or spreading shrub, 0.5-1.5 m high. 
Upper stems and rachises without patent 
hispid hairs, or with patent hispid hairs; short 
curved hairs retrorse or antrorse, sparse to 
dense; stalked glandular hairs absent; sessile 
glands 16-160 mm 2 Cauline leaves 63-104 
mm long, 24-42 mm wide, 2-2.7 times 
longer than wide, base obtuse or broadly 
cuneate (> 60°) or narrowly cuneate (< 60°) or 
attenuate; marginal lobes crenate or serrate, 
regular, 10-21 on each side, acute or obtuse, 
0.9-2.1 mm deep; petioles 16-19 mm long, 
18-25% of lamina length. Lamina upper 
surface indumentum erect, glandular or erect 
or curved, eglandular, 0.25-0.8 mm long, 
sparse (hairs > 0.2 mm apart) or moderately 
dense (hairs 0.1-0.2 mm apart), sessile glands 
16-80 mm 2 ; lower surface indumentum erect, 
glandular or erect or curved, eglandular, 
0.25-0.7 mm long, moderately dense (hairs 
