370 
0.1-0.2 mm apart) or dense (hairs < 0.1 mm 
apart), sessile glands 48-176 mm 2 ; transition 
from leaves to floral bracts abrupt, or gradual. 
Floral bracts elliptical or ovate, 7-30 mm 
long, 3.5-18 mm wide, not consistently 
exceeding verticils. Verticils (inflorescence 
clusters) all separated on rachis, cymes 
entirely monochasial or once dichasial at 
base then monochasial, with 5-14 flowers 
per monochasium, peduncles 0-15 mm long 
on lowermost cluster; bracteoles obovate or 
spathulate or linear, 4.8-7.8 mm long, 0.5-2.5 
mm wide. Corolla tube longer than calyx, or 
same length as calyx; annulus 2.5-3 mm from 
base of corolla, annulus hairs 0.2-0.3 mm 
long; upper lip elliptical, 3.6-47 mm long, 
with eglandular hairs on outer surface; lower 
lip 4-5.8 mm long to end of lateral lobes, 
8-10.5 mm long overall, with 1-20 eglandular 
hairs on platform. Longest stamens 12-14 mm 
long from base of corolla tube; filament hairs 
07-1.2 mm long, mainly at distal end. Style 
12-14 mm long; longer stigma lobe 0.65-0.7 
mm long, shorter stigma lobe 0.4-0.55 mm 
long. Fruiting calyces 0.8-1.5 mm apart on 
rachilla; fruiting calyx cylindrical, 8.2-11 
mm long, 3.4-4.8 mm wide at lobe apices, 
2-2.6 times longer than wide, exterior surface 
with hairs of different sizes or types or with 
all hairs same size and type, hairs glandular 
or hairs eglandular, 0.4-1 mm long, sessile 
glands 80-144 mm 2 ; lobes acute, 2.3-3 mm 
long. Fruiting calyx fringe hairs about the 
same length throughout, 0.15-0.25 mm long 
at apical end, 0.15-0.25 mm long at sinus end, 
sinus hairs absent, inner surface of tube with 
dense ring of long hairs in medial section or 
glabrous. Nutlets 1.9-2.3 mm long. Figs. 5E, 
10G 
Additional specimens examined : Indonesia. Lesser 
Sunda Islands: Timor, undated, herb. Drake (P); Timor, 
Apr 1821, Reinwardt s.n. (L). Australia: Western 
Australia. 8.5 km ENE of Mt Agnes, near headwaters 
on N side of Prince Regent River, Jan 2001, Barrett & 
O ’Connor RLB1588 (PERTH); Secure Bay, side arm on 
W side of bay, Yampi Peninsula, Mar 2001, Barrett & 
Handasyde RLB2047 (PERTH); Ellenbrae Station, Jul 
2004, Brennan & Done 6328 (DNA, PERTH); slopes 
overlooking campsite, Margaret Island, Buccaneer 
Archipelago, Jun 1982, Hopkins BA0475 (PERTH); 
Boongarie Island, 18.3 km NE of Mt Knight, Jun 1988, 
Keighery & Alford 1817 (PERTH); E side of Mindjau 
Creek, Port Warrender, Admiralty Gulf, Jan 1982, 
Kenneally 7764 (PERTH); Campsite on unnamed 
Austrobaileya 9(3): 321-381 (2015) 
tributary of Prince Regent River, arising 19 km SE of the 
mouth, north-west Kimberley, Jun 1984, Kenneally 8922 
(MEL, PERTH); unnamed island in Prince Frederick 
Harbour at entrance to Hunter River, Jun 1984, Kenneally 
8978 (PERTH); junction of unnamed creek and Sale 
River, 30 km ESE of mouth. May 1986, Kenneally 
9603 (MEL, PERTH); Heywood Island (South island), 
Naturalist Island, E side, directly opposite ‘Ninepin’, 
or the Sentinel, Prince Frederick Harbour, May 1987, 
Kenneally 9963 (PERTH); Boomerang Bay, Bigge 
Island, Jun 1972, Marchant 72/60 (PERTH); adjacent 
to creek running out of Koolan townsite. May 1993, 
Mitchell 3112 (PERTH); c. 5 km SW of Crystal Head on E 
side of major river and adjacent to W.A. Water Authority 
gauging station. Mar 1994, Mitchell 3403 (PERTH); 
junction of two creeks where G. Grey had his main base 
camp in 1837, 15° 22’ 04”S 124° 45’ 19”E, Mar 1994, 
Mitchell 3465 (NSW, PERTH); beach on Hanover Bay, 
north Kimberley, May 1998, Mitchell 5401 (PERTH); 
Bachsten Creek, Jul 2001, Riissell-Smith & Handasyde 
TH01-171 (DNA, PERTH); Mallam Spring, SE side of 
Augustus Island, Jul 1990, Willing 187 (PERTH); above 
Moran River Gorge, Prince Regent Nature Reserve, May 
1991, Willing 420 (PERTH); Bonaparte Archipelago, 
May 1972, Wilson 10896 (CANB, PERTH). 
Distribution and habitat : Anisomeles 
principis is found in the Kimberley region of 
Western Australia and on the island of Timor 
(Map 5). In Australia, it reportedly grows 
on sandstone hills and gullies, in Triodia 
hummock grassland, eucalypt woodland or on 
the edges of vine thicket. Two specimen labels 
mention basalt as the substrate. 
Phenology : Flowers collected from January 
to July; fruits from May to July. 
Notes: Anisomeles principis is variable 
with regard to indumentum types present 
on the stems. Several specimens (including 
Kenneally 8922, Wilson 10896) have dense 
antrorse hairs, and one {Kenneally 9603 ) 
has abundant spreading hispid hairs. These 
may prove to be taxonomically distinct. 
Distinguishing features of the species as 
currently circumscribed are: monochasial or 
once-dichasial cymes of the verticils; short 
petioles (16-19 mm); short calyx fringe hairs, 
often pedunculate lower verticils; crowded 
flowers and fruits; and broad bracteoles. 
