398 
typification of that name (Kosteletzky 1835; 
Knaap-van Meeuwen 1970). Kosteletzky 
(1835) quite clearly indicated the single 
typifying elements for both his name and that 
of Linnaeus (1753) and provided accounts of 
both species. Although this does not equate 
to a formal lectotypification in the modern 
sense, subsequent authors have followed 
this citation (e.g. Knaap-van Meeuwen 1970; 
Ding Hou 1996: 606; Ross 1998: 169). Jarvis 
(2007) goes so far as to state “Lectotype 
(Knaap-van Meeuwen in Blumea 18: 23 
(1970): [icon] “ Cynomorium Silvestre ” in 
Rumphius, Herb. Amboin. 1: 167, t. 63. 1741”, 
although Kosteletzky (1835) rather than 
Knaap-van Meeuwen (1970) should probably 
be regarded as making this decision: neither 
state “lectotypify” as such. 
Notes: Cynometra iripa has been described 
as having a style not in line with the dorsal 
suture of the ovary (Knaap-van Meeuwen 
1970), but Australian material has styles 
that are often directly in line with the dorsal 
suture, although during and after anthesis 
they become excentric to varying degrees. 
All living specimens seen in the Cairns area, 
as well as along the Claudie and Olive Rivers, 
have pink ovaries with a dense covering of 
white hairs, thus differing from the rust- 
coloured pubescence described by Ross 
(1998). Fresh specimens from other areas 
were not seen for comparison. 
Sepals have been described as being 
rather long-hairy (Knaap-van Meeuwen 
1970: 23; Ding Hou et al. 1996: 603). Sepals 
of Australian specimens seen by the author 
of this revision do not have a hairy surface 
but do have a ciliate margin especially at the 
apex. 
Etymology: There has been confusion over 
the etymology of the specific epithet iripa. It is 
clear that Rheede’s name for this plant, Iripa , 
was based on an indigenous Malayalam (native 
language of southern India) name (Rheede’s 
opening statement is ‘Iripa Malabarensibus’; 
Ram (2005) also indicates that Malayalam 
names were employed by Rheede). This is 
further confirmed by Nicolson et al. (1988), 
who reported that “Irippa (sic) is still used. It 
is found [in] mangrove swamps, increasingly 
Austrobaileya 9(3): 393-403 (2015) 
scarce in Kerala”. Hence the etymology for 
iripa is derived from Irippa, the Malayalam 
name for the plant. 
2. Cynometra ramiflora L., Sp. PI. 382 
(1753). Type: based on Cynomorium sylvestre 
Rumph., Herb. Amboin. 1: 167, t. 63 (1741), 
fide Knaap-van Meeuwen (1970: 23 “excl. syn 
Iripa Rheede”). 
Cynometra sp. (Paira Homestead Rd 
G.Sankowsky+ 1223); Pedley (2007: 39,2010: 
34). 
Illustrations: Beddome (1869-1874); Pierre 
(1880-1907); Kirtikar & Basu (1918); 
Verdcourt (1979: 84); Whitmore (1983: 255); 
Du Puy (1993: 180 D & E); Corner (1997: 
399); Claussen (2005: 21); Duke (2006: 137) 
as C. iripa (lower photo). 
In Australia: Tree to 15 m, dbh to 60 cm, 
buttressed, blaze red. Bark with numerous 
lenticels, these round or elongated and often 
in vertical lines; twigs with scattered to dense 
round and linear lenticels; new flush foliage 
pink to cream; stipules filiform, c. 1.5 mm 
long; tuft of hairs at petiole apex c. 0.75 mm 
long, caducous. Leaves 1 (rarely 2)-jugate; 
petiole (+ rachis when present) 5-33 mm long, 
shallowly grooved on upper side, glabrous 
on mainland specimens and glabrescent on 
Christmas Island specimens; petiolules 2-6 
mm long, thickened and mostly enclosed by 
leaflet base, glabrous on mainland specimens 
but minutely pubescent on Christmas Island 
specimens; leaflets discolorous, asymmetrical, 
obliquely-ovate, obliquely-oblong, obliquely- 
obovate or broadly elliptical; basal leaflets, if 
present, 17-82 x 9-46 mm; terminal leaflets 
63-210 x 30-98 mm; coriaceous, shiny, 
glabrous; base oblique, rounded, truncate, 
attenuate or cuneate; apex acute, shortly 
acuminate, acuminate or emarginate; margin 
entire; venation brochidodromous, primary 
vein raised on upperside, secondary veins 
in 8-12 pairs (6 -7 pairs on basal leaflets 
if these present), angle of divergence from 
primary vein 40°-50°; tertiary venation 
reticulate. Inflorescences axillary, terminal 
or ramiflorous, solitary or paired (sometimes 
condensed) racemes, up to 20-flowered; rachis 
to 20 mm long, glabrous; pedicels 6.5-12.5 
