6 
3-4 mm long. Calyx 5-lobed; tube 0.25-0.5 
mm long, broadly triangular, lobes spreading 
after anthesis, 0.75-2 mm long, margins 
scarious, glands globular to shortly lineate, 
mostly dark red, occasionally orange. Corolla 
rotate, white to cream to pale pink; tube c. 1 
mm long, lobes spreading after anthesis to 3 
mm long, c. 1 mm wide, apex often weakly 
reflexed, glands dark red often appearing 
black, chiefly lineate, length variable. Stamens 
connivent around the style; base of filaments 
fused to form a rim, attached to corolla-tube, 
uniseriate hairs dense on rim, free portion of 
filament 0.2-1 mm long; anthers 1.5—2 mm 
long, c. 1 mm wide at base, occasional dark 
gland present on connective, apex apiculate. 
Ovary globose, 1-1.5 diameter, tapering into 
style, dark-coloured. Style 2-3 mm long, 
twisted near apex, red-coloured when fresh; 
glands if present, dark red; stigma punctiform. 
Ovules uniseriate, 2-4 embedded in placenta, 
only 1 maturing. Fruit globular to depressed 
globular, 4-7 mm long, 5-7 mm diameter, 
red at maturity, glands dark red and usually 
dense. Seed depressed globular, 4-5 mm long, 
5-6 mm diameter, testa brown, rambling 
spearflower. Fig. 2. 
Additional selected specimens (from c. 118 examined): 
Queensland. Cook District: 1 km SW of the Twin 
Forks, headwaters of the Annan River, Jun 1992, Forster 
PIF10741 et al. (BRI); Home Rule, base of Mt Hedley, 
3 km E of Rossville, Apr 1999, Forster PIF24261 & 
Booth (BRI); Headwaters of Coopers Creek, 3.5 km W 
of Thornton Beach, Jun 1988, Forster PIF4385 & Tucker 
(BRI, CANB, L); Castle Rock, 33.7 km from Mt Carbine 
road on road to Spencer Creek Forestry Camp, Mount 
Windsor Tableland, Whypalla, Nov 1990, Holland 14 
& Hind (NSW); S.F.R. 675, East Mulgrave L.A., Jan 
1977, Gray 269 (BRI); Longlands Gap, S.F. 353, Jun 
1995, Forster PIF16786 (BRI); N.P.R. 904, Parish of 
Palmerston, Feb 1997, Hyland 15588 (CNS); NW of 
Tully, May 2005, Ford 4643 (CNS). North Kennedy 
District: Murray Falls, Murray Upper, Dec 1987, Irvine 
2334 (CNS); Hinchinbrook Island, c. 2 km NW of Mt 
Diamantina, Dec 2000, Kemp TH2582 & Kutt (JCT); 
Burgoo L.A., Garrawalt, Jul 1975, Sanderson 675 (CNS); 
Near Gumburu Environmental School, Paluma, Dec 
2005, Jackes 431 (JCT). 
Distribution andhabitat: Ardisia brevipedata 
is endemic to north-east Queensland where 
it occurs as an understory shrub or small 
tree. It is usually found on the margins or 
in rainforest habitats usually above 500 m 
altitude, although it has been collected in 
Austrobaileya 8(1): 1-23(2009) 
lowland areas. This widespread species 
extends from Paluma in the south ( c . 19°S) to 
the Bloomfield - Windsor Tableland area ( c. 
15°49'S) in the north (Map 2). 
Phenology : Flowers have been collected 
in every month but chiefly from October to 
January. Fruits have been collected throughout 
the year. 
Notes: Hu (1999) considered that Ardisia apus 
Mez from New Guinea was synonymous with 
A. brevipedata ; however, the type description 
of the former refers to gland-like thickish 
points on the margin of the adaxial surface. 
Such glands were absent for all specimens 
of A. brevipedata examined. Unfortunately 
the type specimen of A. apus has been 
lost and according to Sleumer (1988) no 
other collections had been made. The type 
specimen, Ledermann 12015 was collected in 
the Schrader mountains of the Sepik district 
in Papua New Guinea. Hu (1999) indicates 
that collections of A. apus have been recorded 
from the Papua New Guinean provinces 
of East Sepik, Morobe and Milne Bay and 
Hollandia province in Indonesian Papua. 
Etymology: From the Latin brevi - short and 
pedatus - foot, referring to the peduncles 
which are often short. 
3. Ardisia crenata Sims, Bot. Mag. 45: t. 1950 
(1817); Bladhia crenata (Sims) H.Hara, Enum. 
Spermatoph. Jap. Part 1:75 (1948). Type: 
China (Based on Plate 1950, Bot. Mag., from 
material collected/ cultivated by Loddiges 
- Cambridge Botanical Garden). 
Shrub to 1.5(—3) m tall; stem glabrous or 
initially with minute glandular papillae. 
Leaves spirally arranged, petiolate; petiole 
narrowly marginate, 6-10 mm long; lamina 
coriaceous, elliptic-lanceolate, oblanceolate, 
rarely ovate, 6-20 cm long, 2-4 cm wide, 
discolorous, glabrous except for peltate 
scales more common on abaxial surface than 
adaxial surface; apex acute to acuminate, 
base cuneate, margins crenate, undulate and 
revolute, nodules or secretory trichomes 
prominent in the sinuses; midrib raised in a 
shallow groove on adaxial surface, raised on 
abaxial surfaces; lateral veins 12-18 pairs 
on either side of the midrib, looping near 
