Jackes, Ardisia and Tetrardisia 
11 
Etymology : The name refers to the markedly 
crenate margin of the leaves. 
4. Ardisia depauperata (Domin) Jackes status 
nova; Ardisia brevipedata ?var. depauperata 
Domin, Biblioth. Bot. 89: 502 (1928). Type: 
Queensland. CookDistrict: montesBellenden- 
Ker [“Ein Strauch in der mittleren Region des 
Bellenden-ker”], December 1909, K.Domin 
7657 (holo: PR530275 [photograph seen, 
photos at BRI, CANB, CNS and NSW]). 
Ardisia sp. (South Mary LA B.P.Hyland 8778) 
(Bostock & Holland 2007). 
Shrub or small tree 1.5-4 m tall; bark smooth; 
stems often sprawling with adventitious roots, 
branchlets usually zig-zag. Leaves alternate, 
petiolate; petiole 1—3(—4) mm long, reddish, 
weakly marginate, flat on adaxial surface. 
Lamina chartaceous, lanceolate to elliptic, 
often slightly asymmetrical, (4.6-)6.3-9.1(- 
11.2) cm long, 1—2.1(—3) cm wide, adaxial 
surface glossy green, abaxial surface paler, 
glabrous except for scattered scales; apex 
acuminate to obtuse, base cuneate, margins 
smooth, midrib shallowly depressed on 
adaxial surface, raised on abaxial surface; 
lateral veins relatively inconspicuous when 
fresh, more than 15 per side of midrib. Glands 
pellucid when fresh, appearing black when 
dry, from irregularly globular to lineate, 
length:breadth ratio usually less than 3:1. 
Inflorescence axillary, subumbellate to 1 cm 
long; flowers 4-8 per inflorescence; peduncles 
2-4 mm long, reddish; pedicels filiform, 5-10 
mm long, reddish, glands shortly lineate, 
dark red often appearing black; subtending 
bracts soon caducous, 1-1.5 mm long, 0.5-1 
mm wide, glands dark-coloured. Flowers 5- 
merous, 2-2.5 mm long. Calyx 5-lobed; tube 
0.25-0.5 mm long, broadly triangular; lobes 
spreading after anthesis, broadly triangular, 
c. 0.5 mm long, 0.5-0.75 mm wide, margins 
scarious, glands globular to shortly lineate, 
mostly dark red, occasionally some small 
globular, orange glands near margin. Corolla 
rotate, white to cream often with a pink tinge; 
tube c. 0.5 mm long, lobes spreading after 
anthesis to 1.5-2(-2.5) mm long, 1-1.5 mm 
wide, apex often weakly reflexed, glands dark 
red often appearing black, chiefly lineate, 
length variable, occasional small globular 
orange gland near margin. Stamens opposite 
the petals, connivent around the style; base 
of filaments fused to form a rim that is fused 
to corolla-tube, uniseriate hairs present on 
rim, free portion of filament c. 0.2 mm long; 
anthers c. 1.5 mm long and 1 mm wide at base, 
apex apiculate. Ovary globose, 0.75-1 mm 
diameter, tapering into style, dark-coloured. 
Style c. 1.5 mm long, shallow ridges present, 
red when fresh, no glands observed; stigma 
punctiform. Ovules uniseriate, 3 or 4 ovules 
embedded in placenta, only 1 maturing. 
Fruit globular to depressed globular, 5-6 
mm diameter, red at maturity, glands dark 
red, usually dense. Seed globular, 4-5 mm 
diameter, testa brown. Fig.4. 
Additional selected specimens (from c. 23 examined): 
Queensland. Cook District: S.F.R. 144, Fantail L.A. 
Mt Windsor Tableland, Mar 1981, Unwin 757 (CANB, 
CNS, NSW); S.F.R. 144, Mt Windsor, Jul 1976, Unwin 
15 (CNS); Mt Spurgeon, Sep 1936, White 10597 (BRI); 
Root’s Creek, Jan 1936, Flecker s.n. (CNS1984); S.F.R. 
143, South Mary L.A., May 1976, Hyland 8778 (BRI, 
CANB, CNS, NSW); Mt Lewis road, 35-36 km from 
Rex Highway, Carbine L.A., Aug 1986, Weston 656 et 
al. (BRI, CNS, NSW); S.F.R. 143, South Mary L.A., 
Feb 1974, Hyland 7206 (CNS); Mt Lewis road, 35 km 
from junction with Mareeba - Mossman road, Oct 
1987, Foreman 1860 (BRI, CANB, CNS, MEL, NSW); 
Druggies track, Leichhardt L.A., T.R. 66, 11 km along 
Mt Lewis road, Jun 1995, Forster P1F16761 etal. (BRI); 
T.R. 66, Mt Lewis, Aug 1078, Moriarty 2413 (CNS); 
Tinaroo Range, Aug 1971, Moriarty 806 (BRI, CANB); 
Head of Robson Creek, 5.8 km past hoop pine triangle, 
NE end of Tinaroo Dam, Mar 1988, Forster PIF3933 
et al. (BRI, CANB); S.F.R. 185, Edith L.A., Mar 1975, 
Irvine 1254 (BRI, NSW); S.F.R. 194, Barron, Sep 1992, 
Hyland 14567 (CNS, NSW); Mt Bellenden Ker, 1887, 
Sayer s.n. (MEL1612570); Tarzali, Feb 1918, White s.n. 
(BRI [AQ91824], CANB262869). 
Distribution and habitat : This understory 
shrub or small tree has only been collected 
in or associated with rainforest, chiefly at 
altitudes above 800 m. It is endemic to the 
Wet Tropics of North Queensland. Although 
not widely collected it is probably more 
common than collections indicate. It has been 
recorded in areas between the Mount Windsor 
Tableland area (16°12'S) south to the Evelyn 
Tableland area (17°35'S) (Map 4). 
Phenology : Flowering specimens have 
been collected most months of the year but 
particularly in summer and autumn. Fruiting 
specimens chiefly collected from May to 
August. 
