Jackes, Ardisia and Tetrardisia 
17 
combined with the original description and 
notes by C.T.White. Details of the style were 
unavailable because of the nature of the 
specimen. It is the only species seen where 
the corolla lobes are shorter than the tube, 
its taxonomic position thus remains unclear 
until additional fertile material is obtained. 
It differs from both Ardisia brevipedata and 
A. pachyrrhachis in the number of ovules and 
the short thickened peduncle and pedicels, as 
well as the corolla lobes much shorter than the 
tube. 
Etymology : From the Latin fasciculus 
- fascicle or cluster and refers to the flowers 
being clustered in the axils of the subtending 
leaf. 
7. Ardisia pachyrrhachis (F.Muell.) F.M. 
Bailey, Botany Bull., Dept. Agric. Queensland 
3: 14 (1891). Bladhiapachyrrhachis F.Muell., 
Viet. Nat. 8: 15 (1891). Type: Queensland. 
Cook District: In the upper region of Mount 
Bartle Frere, January 1891, S.Johnson s.n. 
(lecto: MEL1612575 [here chosen]; isolecto: 
BRI [AQ23416], K, MEL1612573; P n.v). 
Illustration : Cooper & Cooper (2004: 335). 
Shrub or small tree, 1-5(-7) m tall, sparsely 
branched; bark smooth; branchlets angular, 
reddish-purple when young, scars obvious. 
Leaves alternate, petiolate; petiole (2-)5-10 
mm long, purplish-red, marginate, flat on 
adaxial surface. Lamina coriaceous, obovate, 
7.7-28.5 cm long, (1.7-)2.3-8.3 cm wide, 
adaxial surface dull green, abaxial surface 
paler, glabrous except for scattered scales; 
apex obtuse to acuminate, base cuneate, 
margins smooth, usually recurved, midrib 
depressed on adaxial surface, raised on 
abaxial surface, purplish-red in colour when 
fresh, lateral veins relatively inconspicuous 
particularly when fresh, more than 20 per 
side; glands in reflected light, pellucid when 
fresh, red on oxidation, externally appearing 
black when dry, irregularly globular to 
lineate, with a length:breadth ratio up to 
50:1. Inflorescence axillary, racemose to 
subumbellate or umbellate, to 4 cm long; 
peduncles thick, 5-11 mm long, (1—)2—5 mm 
wide, reddish; rhachis 2-10 mm long, flowers 
(10-)19-44 per inflorescence; pedicels 
filiform, often curved, reddish 10-23 mm 
long, glands appearing black; subtending 
bracts soon caducous, to 10 mm long and 4 
mm wide, scars persistent, glands shortly 
lineate, appearing black. Flowers 5-merous, 
4-5 mm long. Calyx 5-lobed; tube 0.2-03 
mm long, lobes broadly triangular to broadly 
ovate, 0.75-1 x 1-1.5 mm, spreading after 
anthesis, margins scarious, glands dense, 
dark red to black, but near margin they are 
often small, globular and orange-red so calyx 
usually appears almost black when dry but 
dark pink when fresh. Corolla white, cream 
or pale pink; tube c. 0.5 (-1.5) mm long; lobes 
twisted in bud, spreading after anthesis to 
3.5-4 mm long, c. 1.5 mm wide, apex reflexed, 
margins scarious, inner surface with scattered 
uniseriate ferruginous hairs, glands dark 
red, irregularly globular to shortly lineate. 
Stamens opposite the petals, connivent around 
the style; base of filaments fused to form a rim 
fused to corolla-tube, uniseriate ferruginous 
hairs dense on rim, free portion of filament 
to c. 0.3 mm long; anthers cordate-sagittate, 
c. 2 mm long, 1-1.5 mm wide at base, glands 
absent, apex apiculate. Ovary globose, c. 
1.5 mm diameter, tapering into style, pink 
when fresh. Style usually twisted, 2-3 mm 
long, pink, glands dark, stigma punctiform. 
Ovules uniseriate, 4, embedded in placenta. 
Fruit globular to depressed-globular, 5-8 
mm diameter, bright red at maturity, style 
persistent, glands shortly lineate dark- 
coloured. Seed depressed-globular, 4-5 x 6 
mm, testa brown, mountain Ardisia. Fig. 7. 
Additional selected specimens (from c. 107 examined): 
Queensland. Cook District: Upper Parrot Creek, Annan 
River, Sep 1948, Brass 20239 (BRI, CANB); Gap Creek, 
c. 38 km S of Cooktown (9 km by road from Rossville), 
Sep 1960, Smith 11231 (BRI); T.R. 146, Tableland L.A., 
Sep 1980, Hyland 10583 (CNS); Mt Misery on Mt 
Carbine Tableland, Sep 1972, Webb & Tracey 10823 
(BRI, K); Mt Hemmant just N of Noah Creek in Cape 
Tribulation area, Jul 1973, Webb & Tracey 11726 (BRI); 
S.F.R. 144 Whypalla, Bowerbird L. A., Feb 1988, Hyland 
13506 (CNS); S.F.R. 144, Western edge of Windsor 
Tableland, Nov 1971, Dockril! 301 (BRI, CNS); S.F.R. 
143 Kanawarra, Carbine T.R., May 1995, Gray 5936 
(CNS); Daintree N.P., near end of Mt Lewis road, 12 km 
SW of Mossman, Nov 1988, Jessup GJM143 et al. (BRI); 
Mt Lewis road, 28-29 km from Rex Highway, Aug 1986, 
Weston 602 et al. (BRI, NSW); 8 km along Mt Lewis 
road from the junction with Mareeba to Mossman road, 
Oct 1987, Foreman 1676 (AD, BRI, CANB, CNS, MEL, 
NSW); T.R. 66, Mt Lewis, Sep 1978, Moriarty2465 (BRI, 
CNS); Murray Prior Range, tributary of Hills Creek, 
