Jackes, Ardisia and Tetrardisia 
21 
Gosschalk L.A., Mar 1995, Gray 6020 (CNS); S.F.R. 755 
Bartle Frere, Gosschalk L.A., Nov 1991, Hyland 14340 
(BRI, CNS); Near Topaz, Feb 1990, Nicholson s.n. (CNS 
93290); N.R 226, Bartle Frere zig-zag track from Russell 
River to Towalla, 250 m from river, Sep 1993, Bostock 
1467 & Turpin , (BRI); Junction track, Russell River, Apr 
1948, Brass 18261 (BRI). 
Distribution and habitat : Most collections 
have been made in rainforest associated 
with Mt Bartle Frere and Mt Bellenden Ker 
at altitudes above 600 m. However, several 
specimens of Tetrardisia bifaria have been 
collected at lower altitudes such as at The 
Boulders, Babinda (alt. 40 m) and at 220 m in 
the upper Goldsborough Valley. Although this 
species has a restricted distribution it chiefly 
occurs within protected areas (Map 7). 
Phenology: Flowers have been collected 
from August to April with a peak occurring 
in January and February. Fruits recorded 
from April to November but mature fruits 
have been chiefly observed in September and 
October. 
Notes: The species is readily distinguished 
from the Australian Ardisia species by the 4- 
merous flowers and the sessile or subsessile, 
2-ranked leaves where the cordate lobes of the 
leaf base appear to clasp the stem. The flowers 
and fruits are often difficult to see as they are 
concealed under the leaves. 
Etymology: The specific epithet is from the 
Latin hi- two and -farius -ranked, referring 
to the leaves being two ranked. 
Excluded names 
Ardisia pseudojambosa F.Muell., Fragm. 4: 
81 (1864) [=Tapeinosperma pseudojambosa 
(F.Muell.) Mez, fide Jackes (2005)]. 
Ardisia repandula F.Muell., Fragm. 4: 
82 (1864) [=Tapeinosperma repandulum 
(F.Muell.) Jackes, fide Jackes (2005)]. 
Acknowledgements 
I would like to thank the Directors of the 
following herbaria - BRI, CANB, MEL, 
NSW and QRS (now incorporated at CNS) 
for the loan of specimens: Dr O.Sida (PR) 
for providing the type photograph of Ardisia 
depauperata , Dr Paul Forster and staff of 
the Queensland Herbarium for advice and 
technical assistance. 
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