Booth et al ., Four new Cyperus 
Table 2. Correlations of character states and ordination vectors. 
39 
Character state 
Correlation 
Culm cross section trigonous 
0.9668 
Culm cross section triquetrous 
0.9668 
Base of primary inflorescence bract teeth absent 
0.9528 
Base of primary inflorescence bract teeth present 
0.9528 
Involucral bracts septate nodules absent 
0.9151 
Involucral bracts septate nodules present 
0.9151 
Style length (mm) 
0.8489 
Glume mucro length <0.3 mm 
0.8084 
Glume mucro length 0.3-0.7 mm 
0.8084 
Rachilla wing absent 
0.7224 
Rachilla wing present 
0.7224 
wide, with 1-7 spikelets. Spikelets 5-25 mm 
long, 2.5-3.2 mm wide, 10-40(-50)-flowered, 
digitate, spreading, linear, reddish brown to 
dark brown to dark grey to black. Glume s 
distichous, 2.5-3.5 mm long, dark reddish 
or red-brown or blackish-red or ferrugineous 
or streaked red-brown, apex mucronulate or 
mucronate or acuminate, mucro excurved or 
straight, one third to two thirds imbricate, 
keeled, 1.1-1.3 mm apart, 4-10 nerved, with 
arcuate keel. Rachilla straight, persistent, 
winged. Stamens 3; anthers linear; connective 
setulose; 0.5-0.9 mm long. Style shorter than 
stigmas, 3-fid, 1.2-1.5 mm long. Nut obovoid 
to ellipsoid; trigonous to triquetrous, or 
dorsiventrally compressed or flattened with a 
flat or concave face against the rachilla, half 
as long as the glume, 1.5-2 mm long, 1-1.25 
mm wide, smooth or densely granulose or 
granulate, with sides exposed, apex apiculate 
or rounded, brown to golden brown. 
Additional selected specimens examined : Papua New 
Guinea. Eastern Highlands Province: Kini Creek, NE 
slopes of Mt Michael, Sep 1959, Womersley NGF11715 
(BRI). Morobe Province: 5 km from Markham River 
Bridge along road to Labu, Jul 1992, Forster PIF11067 
& Liddle (BRI, LAE). Northern Province: N slope of 
Mt Lamington, Jul 1953, Hoogland3320 (BRI). Central 
Province: E slope of Lake MyolaNo. 2, Sep 1973, Croft 
NGF34536 (BRI). Queensland. Cook District: Davies 
Creek, S.F.607,9 km pastN.P. carpark, Jun 1991, Forster 
PIF8536 (BRI); Bellenden Ker, 40 m below tower 3, May 
2001, Booth et al. 2614 (BRI, K, NSW); Foothills of Mt 
Bellenden Ker, May 2001, Cooper 1529 (BRI); Bellenden 
Ker Cable Car Station, Jan 2002, Booth et al. 2834 
(BRI); Junction Creek, Russell River, Apr 1948, Brass 
18262 (BRI); Wooroonooran N.P., CSIRO EP/34, off 
Russell River track, Jan 2007, Ford AF4910 & Metcalfe 
(BRI, NSW); Bartle Frere, Oct 1935, Blake 9758 (BRI); 
East Bartle Frere, Nov 1994, Hunter JH1837 (BRI). 
Wide Bay District: Fraser Island, Oct 1930, Hubbard 
4396 (BRI); Fraser Island, May 1964, Webb 6328 (BRI, 
CANB); track up to Mt Bowarrady tower, Fraser Island, 
Sep 1986, Russell-Smith 1809 & Lucas (BRI). Moreton 
District: McDonald Road, 3 km N of Peachester, Jul 
1993, Bean 6256 (BRI); Mooloolah Scrubs, Dec 1890, 
Field Naturalists s.n. (BRI [AQ 647235]); Base of 
Buderim Mt., Mar 1934, Blake 5251 (BRI). 
Distribution and habitat : Cyperus 
pedunculosus is found throughout coastal 
Queensland from Buderim north to Davies 
Creek near Cairns. It is also widely distributed 
in New Guinea. Plants occur in wet forests 
often along shady creek banks. 
Notes : A number of varieties have been 
named under Cyperus pedunculosus , three 
from Papua New Guinea (Kiikenthal 1924, 
1939, 1943) and one from north Queensland 
(Domin 1928). Blake (1947) and Kern (1974) 
were of the opinion that these varieties and 
also Cyperus montis-sellae K.Schum. were 
merely minor variations of C. pedunculosus 
and this is also our conclusion based upon 
the protologue descriptions by Schumann 
