Editorial Board 
Dr Shane Ahyong 
Dr Don Colgan 
Dr Elena Kupriyanova 
Dr Andrew Mitchell 
Dr Robin Torrence 
Editor 
Dr Shane McEvey 
Journal compilation © 2016 Australian Museum, Sydney 
No part of this publication may be reproduced without 
permission of the Editor. 
Volume 68 Number 1 
Published (print and online) 10 May 2016 
Price: AU$40.00 
Printed byRodenprint Pty Ltd, Sydney 
ISSN 0067-1975 (print) 
ISSN 2201-4349 (online) 
The Australian Museum is a statutory authority of, 
and principally funded by, the NSW State Government. 
NSW 
GOVERNMENT 
The Australian Museum houses some of the world’s most 
important collections of Australian animal, fossil and geological 
specimens and cultural objects. Research on these millions 
of specimens and artefacts yields insights into how our world 
changes through time and how its diversity can be classified, 
interpreted, and appreciated. This knowledge, when shared 
among the scientific and broader community—initially through 
publication — helps us understand the significance of the impact 
we have on our environment. Furthermore, we come to know 
with more certainty what reasonable steps society can take now 
for the well-being of future generations. Our responsibility is 
also to inspire the exploration of nature and cultures; our vision 
is a beautiful and sustainable natural world with vibrant and 
diverse cultures that we are able to see and know. 
Since 1889 the results of studies on Australian Museum 
collections, or studies that more generally lead to a better 
understanding of nature and cultures in Australia and the Pacific, 
have been published by the Museum in its premier science 
journal Records of the Australian Museum. In 1999 we began 
releasing PDF of published articles through our open access 
website. In 2008 we adopted DOI registration for our online 
content to facilitate persistence and cross-linking in the scientific 
literature. In 2009 we digitized the entire legacy of all science 
published by us since 1851, and made that huge searchable 
resource permanently and freely available online. To accelerate 
publication of peer-reviewed science we are adopting a one- or 
several-article per publication model from volume 65 (2014) 
and we are limiting, but not abandoning, print production. 
There were six issues in 2015. All that is published in print is 
immediately and freely available online. 
Authors are invited to submit manuscripts to the Editor. 
Manuscripts meeting subject and stylistic requirements outlined 
in the Instructions to Authors (see inside back cover) are peer- 
reviewed by external referees to meet standards of excellence 
set by the Editorial Board. 
http://dx.doi.org/issn.2201-4349 
Records of the Australian Museum is covered in the Thomson 
Reuters Scientific services: Current Contents® / Agriculture, 
Biology, and Environmental Sciences, and Science Citation 
Index Expanded (also known as SciSearch®) 
We encourage cross-linking in the scientific literature by applying 
DOI registration to Australian Museum publications via CrossRef ® 
Bibliographic information is openly available atMendeley.com 
