Sinclair: Hydropeza empidid flies of Australia 
15 
Hydropeza intricata sp. nov. 
Figs 4, 20, 24 
Type material. Holotype 8, labelled: “AUST: N.QLD:/ 
14.7km/ up Mt. Lewis Rd., 860m/ 22.iv. 1994, cascading/ 
creek, B. J. Sinclair/ 16°30'S 145 0 17'E”; “HOLOTYPE/ 
Hydropeza/ intricata/ Sinclair [red label]” (AMS). Paratypes: 
Queensland: 2 88, 4^$, same data as holotype (AMS); 
388 , 1?, 31.6 km up Mt. Lewis Rd., 1000 m, 21.iv. 1994, 
cascading ck, 16°30'S 145°17'E, B. J. Sinclair (CNC). 
Diagnosis. This species is distinguished from other 
Queensland species by acrostichal setae extending to 
between first and second dorsocentral setae; male scutellum 
with 1 pair of elongate discal setae; male mid femur with 
row of short basal posteroventral setae and mid tibia with 
short row of spine-like setae on apical third; male hind 
trochanter with 2 spine-like anteroventral setae; and male 
cercus unmodified and apex of the phallus is inflated and 
highly complex. Females are distinguished from all other 
species by 1 pair of discal setae on the scutellum and segment 
8 broad and inflated, not flattened. 
Description. Head and abdomen dark brown, thorax 
brown with paler patches; coxae brownish yellow, legs 
progressively darker towards tips. Wing length 3-3.3 mm. 
Male. Head. Ocellar setae very long, erect, divergent, inserted 
posterior to anterior ocellus; ocellar tubercle with some long 
setulae; postocellar seta stout and dark; upper postocular 
setae stout and dark, lower setae fine and slender; postgenal 
setae similar to lower postoculars. Face without setulae. 
Scape and pedicel slightly paler than postpedicel; scape short 
and slender, slightly longer than length of globular pedicel; 
postpedicel not greatly prolonged, 3x longer than scape, 
basal portion elongate-oval, slightly longer than slender, 
straight apical portion; arista-like stylus distinctly longer 
than postpedicel, concolorous with postpedicel. Proboscis 
stout and robust, extended beyond mid-length of fore coxa, 
directed posteriorly; palpus half-length of proboscis, clothed 
in long pale and dark setae; apex of palpus rounded; apical 
third of palpus somewhat flattened with dense pubescence. 
Thorax. Clothed in fine pruinescence; mesonotum brown, 
with postpronotal lobe, lateral margin and postalar ridge pale 
brown; pleura paler than notum, pale along sutures. Setae 
generally long and stout; 2-3 acrostichal setulae extending 
to between 1st and 2nd dorsocentral setae, less than 0.5x 
length of dc; pprn with 1 seta and several short setulae; 1 
long presut spal; 5 uniserial dc, about subequal in length 
with several setae interspersed; 3 npl; 1 long psut spal and 
1-2 finer setae; 1 pal seta; 1 pair of sctl, with 1 pair of finer 
outer marginal setae; disc of scutellum with 1 pair of long 
setae. Antepronotum with pair of pale setulae. 
Legs. Coxae and femora ventrally yellowish-brown; 
remaining segments increasingly darker apically. Inner 
anterior margin of fore coxa with some 15-20 dark, 
variously lengthened and thickened spine-like setae, mostly 
concentrated apically; basal most seta longer than width 
of coxa. Anterior surface of mid and hind coxae with long 
brown setae. Legs clothed in very long setae as typical for 
genus, especially ventral faces. Fore femur straight, inflated 
(subequal in thickness to mid femur), with row of 2 stout 
anterior setae at apical fifth; 5-7 posterodorsal setae, apical 
2 setae distinctly stouter. Fore tibia with 3 anteroventral 
setae on apical third; 2 anterodorsal and 2 posterodorsal 
setae; 1 anterodorsal preapical seta. Fore tarsomere 1 with 
1 basal anteroventral and 1 preapical anterovental seta, more 
than 2x width of segment; 1 long basal ventral, 2 shorter 
ventral setae at middle and apex. Mid femur with row of 
3-4 anterodorsal setae stouter towards apex; 1 anterior and 
1 posterior seta on apical third; basal half with row of 4-5 
short, stout posteroventral setae, less than width of femur; 
apex with 1 anterior, 1 dorsal and 1 posterior preapical seta. 
Mid tibia with width reduced on ventral apical third, with 
row of 5-8 short spine-like setae; apex with short spur-like 
ventral seta; 2 posterodorsal setae; apex with 1 dorsal seta. 
Mid tarsomere 1 with 1 long anteroventral basal seta. Hind 
femur inflated, nearly 1.5x broader than fore or mid femora, 
without erect dorsal setae; with anteroventral row of stout 
setae along entire length; apical fourth with 2 anterodorsal, 1 
posterodorsal, 1 anterior and 1 posterior seta. Hind trochanter 
with 2 spine-like anteroventral setae. Hind tibia with 2 
anterodorsal, 1 anteroventral and 4-5 posterodorsal setae; 
1 preapical anterodorsal and 1 apical anteroventral seta; 
apical fourth with dense posterior setae. Hind tarsomere 1 
without erect setae. Fore and mid tarsomeres slightly longer 
than tibia; ventral apical margin of tarsomere 4 of fore and 
midlegs somewhat pale, flattened and expanded; tarsomere 
4 of hindleg not laterally compressed ventrally; tarsomere 
5 of each leg lacking dorsoapical extension. 
Wing. Infuscate with 1 long basal costal seta, longer than 
sctl; all veins lacking setulae; R 4 and R 5 distinctly divergent 
apically; bm-m complete; cell dm slightly produced, longer 
than cell bm; CuA + CuP reduced to streak. Halter brown. 
Abdomen. Tergites and sternites clothed in setae; setae 
of posterior margin overlapping more than half length of 
following tergite. T8 broad, nearly one-half length of T7; 
posterior marginal setae slender, some twice length of tergite. 
Terminalia (Fig. 24). Cercus projected horizontally, 
shorter than epandrium, not forked or subdivided; apex 
sharply pointed ventrally. Epandrium broad with scattered 
setae. Surstylus slightly tapered, apex broadly rounded, not 
arched medially. Hypandrium subrectangular broad; apical 
margin shallowly notched; gonocoxal apodeme broad, 
rounded; postgonite absent. Phallus broad with complex 
apical half, inflated with 2 pairs of elongate, anteriorly 
projecting processes; apex of phallus arrowhead-shaped, 
projecting posteriorly; ejaculatory apodeme L-shaped, longer 
than gonocoxal apodeme. 
Female. Similar to male except as follows: scutellum 
with 1 pair of discal setae; mid femur lacking row of stout 
posteroventral basal setae; mid tibia not narrowed apically, 
lacking row of spine-like setae; hind femur not strongly 
inflated, lacking anteroventral row of stout setae. Terminalia 
(not dissected): segment 8 inflated, broad; cercus slightly 
longer than epiproct, nearly 2x longer than wide; broad 
with rounded apex. 
Distribution. This species is recorded from above 800 m on 
Mt. Lewis, Queensland (Fig. 20). 
Etymology. The specific name is from the Latin intricatus 
(entangled, complicated), in reference to highly complex 
and unusual apex of the phallus. 
Remarks. Female segment 8 is apparently inflated and not 
flattened as in related species. It is uncertain whether this 
is an artefact of drying, but other distinguishing features 
(e.g., acrostichals and scutellar setae) are consistent among 
specimens. 
