Editorial Board 
Dr Shane Ahyong 
Dr Don Colgan 
Dr Elena Kupriyanova 
Dr Andrew Mitchell 
Dr Robin Torrence 
Dr Ross Sadlier 
Editor 
Dr Shane McEvey 
Journal compilation © 2014 Australian Museum, Sydney 
No part of this publication may be reproduced without 
permission of the Editor. 
Volume 66 Number 4 
Published (print and online) 28 May 2014 
Price: AU$30.00 
Printed byRodenprint Pty Ltd, Sydney 
ISSN 0067-1975 (print) 
ISSN 2201-4349 (online) 
The Australian Museum is a statutory authority of, 
and principally funded by, the NSW State Government. 
NSW 
GOVERNMENT 
The Australian Museum houses some of the world’s most 
important collections of Australian animal, fossil and geological 
specimens and cultural objects. Research on these millions of 
specimens and artefacts yields insights into how our world changes 
through time and how its diversity can be classified, interpreted, 
and appreciated. This knowledge, when shared among the scientific 
and broader community—initially through publication—helps 
us understand human impact on our environment and what 
reasonable steps society can take now for the well-being of future 
generations. Our responsibility is to inspire the exploration of 
nature and cultures; our vision is a beautiful and sustainable 
natural world with vibrant and diverse cultures. 
Since 1889 the results of studies on Australian Museum 
collections, or studies that more generally lead to a better 
understanding of nature and cultures in the Australian Region, 
have been published by the Museum in its premier science journal 
Records of the Australian Museum. In 1999 we began releasing PDF 
of published articles through our open archive website. In 2008 
we adopted DOI registration for our online content to facilitate 
permanence. In 2009 we digitized the entire legacy of all science 
published by us since 1851, and made that huge searchable 
resource permanently and freely available at our website. To 
accelerate publication of peer-reviewed science we are adopting 
a one- or several-article per publication model from volume 65 
and we are limiting, but not abandoning, print production. The 
number of issues per volume will, from 2014, not be limited 
to three. All that is published in print is immediately and freely 
available online. 
Authors are invited to submit manuscripts to the Editor. 
Manuscripts meeting subject and stylistic requirements outlined 
in the Instructions to Authors are peer-reviewed by external referees 
to meet standards of excellence set by the Editorial Board. 
http://australianmuseum.net.au/ Scientific-Publications 
Search publications at 
http: //australianmuseum. net. au/ j ournalfinder 
Records of the Australian Museum is covered in the Thomson 
Reuters Scientific services: Current Contents® / Agriculture, 
Biology, and Environmental Sciences, and Science Citation Index 
Expanded (also known as SciSearch®) 
We encourage cross-linking in the scientific literature by applying 
DOI registration to Australian Museum publications via CrossRef ® 
