Lonsdale: Australian Heteromeringia flies (Clusiidae) 
237 
Species descriptions (alphabetical order) 
Heteromeringia asteia n.sp. 
Figs 17-21, 88 
Type material. Holotype: Tasmania: Hellyer Gorge, 41°16'24"S 
145°36'55"E, on horizontal log, 20.xii.2003, S.A. Marshall (1$, AMS). 
Paratypes: Australian Capital Territory. 35.22S 148.50E, Blundells Ck., 
D.H. Colless, i.1988 (2$S, ANIC; 2$$ USNM),ii.l987 (1?, ANIC). New 
South Wales: Coombadjah Ck., Washpool S.F., lO.i. 1982, BJ. Day (\S, 
AMS), Carrai SF, 30°58’48"S 152°17’06"E, 975 m, E. Tasker, 3-8.xii.1997, 
sticky trap on E. viminalis , CS-RO-127-1 (1$, AMS), Carrai SF, 30°59’45"S 
152° 16’23"E, 930 m, E. Tasker, 3-8.xii. 1997, sticky trap on E. campanulata, 
CS-FZ-127-6 (1 $, DEBU), Werrikimbe NP, 31°H'56"S 152°10'23"E, 1025 
m, E. Tasker, l-7.xii.1997, sticky trap on E. campanulata, WC-WN-127-1 
(1$, AMS). Queensland: N QLD, Birthday Crk. near Paluma, 183.1967, 
D.K. McAlpine & G. Holloway (1$, AMS). Tasmania: Hellyer Gorge, 
41°16’24"S 145°36’55"E, on horizontal log, 20.xii.2003, S.A. Marshall 
(Id 1 , DEBU), 42.44S 146.25E, 4km SW by W, T. Shea, 460 m, 3.ii. 1980, 
Lawrence & Weir (1$, ANIC), 42.13S 146.01E, Fra nkli n R„ 223.1983, 
I.D. Naumann & J.C. Cardale, ex. ethanol (1$, ANIC). 
Description 
Male (Fig. 17). Body length 3.3-5.0 mm. Anepisternal disc 
present and relatively large, multiple times width of base 
of anepisternal bristle. First flagellomere orbicular. Bristles 
black. Arista closely pubescent. Vibrissa relatively long and 
curved. Ocellar bristle thin and short, but distinctly longer 
than ocellar tubercle. Two dorsocentral bristles. Gena shallow 
and not strongly angled or bent. Face and buccal cavity 
pilose and meeting at an angle. Posterior Vi- 2 A of frons dark 
brown (lateral and posterior margins paler) and anterior 2 /3 
tomentose; back of head, occiput (sometimes only posterior 
margin) and posteroventral corner of gena dark brown; 
antenna (excluding arista) light yellow (Tasmanian specimens 
with brown infuscation bordering base of arista on inner face); 
face with dark yellow to orange tint; clypeus dark brown; 
remainder of head yellow; upper % of gena pilose. Thorax 
dark brown with postpronotum dark piceous, notopleuron 
slightly lighter brown, lateral margins of scutellum and 
posterolateral corner of scutum light yellow and ventral 
margin of katepisternum yellow; scutum sometimes with thin 
yellowish stripes extending from lateral half of anatergite to 
side of scutellum and anterior dorsocentral. Legs yellow with 
fore tarsi, fore tibia and inner-distal spot on fore femur dark 
brown; Tasmanian specimens with mid tibia light brown sub- 
basally, hind tibia dark brown sub-basally and hind femur with 
light apical infuscation on outer face. Abdomen dark brown. 
M 1+2 ratio 5.5-8.0. Wing dark on distal Ms (fading posteriorly) 
and with cloud around cells br and dm (infuscation sometimes 
restricted to region immediately around veins). Halter white 
with base and side of stalk infuscated. 
Female. As described for male except as follows: scutum and 
pleuron entirely dark brown (scutellum and anatergite still 
with yellowish spot); first flagellomere with anterior margin 
and dorsal % of inner face (or entire inner face) brown; face 
orange; ventral margin or medial longitudinal stripe on gena 
brownish; frons black with anterior margin and posterolateral 
corner orange; anterior half of frons tomentose. NSW 
females smallest of material examined, with distal l A of fore 
femur and base of hind tibia dark brown. ACT and Tasmanian 
females somet im es with palpus, first flagellomere, face and 
ventral half of gena darker. 
Male terminalia (Figs 18-21). Epandrium as long as high 
and wider than long. Surstylus small and rounded with 
several inner-distal tubercle-like bristles. Length of cerci 
half width of epandrium, widest medially and emarginate. 
Hypandrium subtriangular, with bristles along basal and 
posterior margins; with one small anterior lobe bearing two 
stout bristles. Distiphallus twisted medially with fan-shaped 
frill and two distal/medial U-shaped sclerites. 
Female terminalia (Fig. 88). Spermatheca dark with 
apex barely invaginated; width approximately % length. 
Spermathecal duct not longer than twice length of 
spermatheca. Ventral receptacle small with loosely coiled 
and relatively short flagellum. 
Etymology. The specific name is derived from a Greek word 
for “pretty” ( asteios ). 
Comments. This attractive and widespread species is 
characterized by a large body size, yellowish notal patches 
and a distinct wing pattern. It is similar to Heteromeringia 
australiae in having a large male anepisternal disc, minute 
ocellar bristles and a strong medial wing spot, but the latter 
has confluent spots on the wing, (sometimes) pale patterning 
on the notum and femora, distinct male genitalia and shorter 
spermathecae. 
Heteromeringia australiae Malloch, 1926 
Figs 7, 8, 15, 22-27, 89-91 
Heteromeringia australiae Malloch, 1926: 48. Frey, 1960: 
25. D.K. McAlpine, 1960: 73. Sasakawa, 1966: 79. 
Type material. Holotype: New South Wales: North Coast, Coramba 
(1 (j, AMS), [not examined]. 
Material examined. Australian Capital Territory: Black Mtn., xii. 1987, 
M. Irwin, Malaise trap (1 DEBU). New South Wales: Terania Ck. Near 
Lismore, D.K. McAlpine & K.C. Khoo, 5.ii.l983 (1^, AMS; \S USNM), 
6.ii. 1983 AMS), 4 mis SW of Jervis Bay, “8.10.1949”, K.H.L. Key 
(1$, ANIC). Queensland: N QLD, The Crater, near Herberton, D.K. 
McAlpine & G.A. Holloway, 293.1972 (1$, USNM), 303.1972 (1$, 
AMS), Mt. Haig 2km NE by E of Atherton, 18.xi.1981, D.H. Colless, 
Malaise trap (Id 1 , ANIC), 17.27S 145.29E, Hugh Nelson Range, l.viii-1. 
ix.1995, L. Umback, 1150 m. Malaise trap (Id, DEBU), 17.06S 145.37E, 
GS2, Mt. Edith, 31.v-30.vii.1995, P. Zborowski, Malaise trap (Id, ANIC), 
Birthday Ck. Falls via Paluma, 11-12.V.1980, I.D. Nauann & J.C. Cardale 
(1 $, ANIC), Danbulla For. Res., 13kmNEbyNofYungaburra, 17.xi.1981, 
D.H. Colless, Malaise trap (1$, ANIC). Tasmania: 41.50S 146.03E, Pelion 
Hut, 3km S Mt. Oakleigh, 860 m, 83-12.ii.1991, A. Calder, W. Dressier, 
malaise #5, closed forest (Id, ANIC). 
Redescription 
Male (Figs 7, 22). Body length 4.1-4.5 mm. Anepisternal 
disc present and very large (approximately 2-3 times wider 
than that of other species). First flagellomere orbicular. 
Bristles black. Arista pubescent. Vibrissa relatively long and 
curved. Ocellar bristle thin and half length of postvertical. 
Genal bristles relatively long. Two dorsocentral bristles. 
Gena high and sometimes angled at midpoint. Face and 
buccal cavity evenly curving and velvety. Frons dark brown 
with lateral margin yellow and posterior margin (enclosing 
brown ocellar tubercle) reddish; first flagellomere with distal 
% of inner face (sometimes faded dorsally) and distal V 3 -V 2 
of outer face (sometimes variably faded) infuscated; back 
