Lonsdale: Australian Heteromeringia flies (Clusiidae) 
239 
of head dark brown above foramen; dorsal half of gena light 
yellow and silvery tomentose and ventral half of gena dark 
brown to orange/dark yellow; palpus light brown (paler to 
base); anterior margin of occiput light yellow or entirely 
pale; face and buccal cavity reddish; anterior half of frons 
pilose. Scutum grey-yellow (orange to sides) with anterior 
margin (including anterior half of postpronotum, which is 
otherwise rusty) brown and with central stripe continuing 
onto yellow scutellum; also with one pair of lateral floating 
brownish stripes. Laterotergites dark brown. Pleuron dark 
brown with posterior half of meron yellow. Fore legs with 
entirely dark brown; anterior half of mid coxa dark brown; 
base of mid femur light brown or basal half brown; hind 
femur brown apically and (sometimes) medially; hind tibia 
dark brown on basal l A, lightest subapically; remainder 
of legs yellow. Fore tarsi slightly compressed laterally. 
Abdomen dark brown. M 1+2 ratio 3.7-4.0. Wing dark with 
clear subapical band open to posterior margin of wing and 
with anterobasal margin lighter. Halter white with base and 
side of stalk infuscated. 
Female. As described for male except as follows: thorax dark 
brown with postpronotum reddish and scutum with one pair 
of thin yellowish stripes continuing onto sides of scutellum; 
legs yellow with fore tibia and tarsi dark brown and fore 
femur with large inner-distal spot and small outer-distal spot; 
first flagellomere only infuscated along anterior margin; gena 
and parafacial yellowish-orange; apical margin of palpus 
very weakly infuscated; occiput yellow; terminalia yellow. 
Male terminalia (Figs 24, 26, 27). Epandrium relatively 
small. Cerci large and united, widest medially and emarginate 
apically. Surstylus thin and pointed, sparsely setulose on 
outer face and with several inner-distal tubercle-like bristles. 
Hypandrium+pregonite with two subequal distal lobes 
(anterior lobe setulose on inner and outer faces, and posterior 
lobe with apical bristle behind long shallow carina). Ribs 
of distiphallus of nearly equal length, with one terminating 
in reticulated leaf-like structure; both ribs with recurved 
medial processes. 
Female terminalia (Fig. 89). Ventral receptacle short and 
rounded with flagellum apical, very long and straight. 
Spermatheca smooth, as long as wide (at base), slightly 
tapering apically and with several basal wrinkles. 
Comments. Aside from the females described above, there 
are several others belonging to what are possibly one or 
two undescribed species. They differ in that the face meets 
the buccal cavity at an angle, the halter is light brown, 
the palpus is dark brown either apically (NSW—genitalia 
Fig. 90) or basally (QLD—genitalia Fig. 91), the wing 
infuscation extends to surround A!+CuA 2 basally (Fig. 
8), the notum is almost or completely dark brown and the 
internal genitalia differ slightly in morphology, although this 
latter difference may be negligible. One of these females 
may be the counterpart to the smaller male outlined below, 
but it will not be described as distinct until more material 
is available. [Label data: NSW: Bruxner Park FR, N Coffs 
Hbr., 30°15'S 153°06'E, 200 m, yellow pans on trib. Bucca 
Bucca Ck., 16.x. 1999, D. Bickel (1?$, AMS), Werrikimbe 
NP, 31°1F56"S 152°10'23"E, 1025 m, E. Tasker, 29.i-4. 
ii. 1998, sticky trap on E. camaronii, WC-WN-018-2 (1$, 
AMS). QLD: E of Mt. Edith, 800 m, r’for ck, Malaise, B. 
Sinclair, 22-27.iv.1994, 17.05S 145.38E (1$, AMS), N 
QLD, Mt. Edith Forest Road, 1.5 mi off Danbulla Road, 
6.v. 1967, D.H. Colless (3??, ANIC), Danbulla For. Res. 
13km NE by N of Yungaburra, 17.xi. 1981, Malaise trap, 
D.H. Colless (15, ANIC)]. 
In addition to the more widespread “typical” male 
described above, there is a smaller male with slightly 
different colouration and genitalia (Figs 23, 26) that may 
represent a distinct species. Label data: N QLD, The Crater, 
near Herberton, 294.1972, D.K. McAlpine & G.A. Holloway 
(1$, AMS). This male can be diagnosed using the following 
couplet: 
1 Body length 4.1-4.5 mm. Scutum dark brown with pale shoulders, 
or orange to grey-yellow in ground colour with dark medial and 
sublateral stripes. Fore coxa and femur entirely dark brown. Mid 
coxa dark brown on anterior half. Basal l A-Vi of mid femur light 
brown to dark brown. Hind tibia dark brown at base and brownish 
subapically. Gena yellow with venter dark brown. Surstylus 
straight. Medial lobe on phallus broadly rounded and with recurved 
process; without bilobed anterior process. typical male 
-Body length 3.4 mm. Scutum dark brown with shoulders reddish. 
Fore coxa and femur yellow with dark inner-distal spot on femur. 
Mid coxa, mid femur and hind tibia entirely yellow. Gena yellow. 
Surstylus with posterior curve. Medial lobe on phallus short and 
serrate; anteriorly with bilobed process 
smaller male 
