Lonsdale: Australian Heteromeringia flies (Clusiidae) 
241 
Heteromeringia bisetosa n.sp. 
Figs 12, 28-31,92 
Type material. Holotype Australia, Queensland: 15.30S 145.16E, 
lkm SE of Mt. Cook, 14.x. 1980, D.H. Colless, Malaise trap (13, ANIC). 
Paratypes Australia, Queensland: 12.43S 143.17E, 9km ENE of Mt. 
Tozer, 5-10.vii.1986, D.H. Colless (1$, ANIC), 12.44S 143.14E, 3km 
ENE Mt. Tozer, 28.vi-4.vii.1986, D.M. Colless (13, DEBU). Papua New 
Guinea, Nadzab, Markham, R. val., N Guinea, 12.vii.1944, K.V. Krombein, 
E. forkNgafir, cr. 1000-2000ft., native trail, ex. colln K.V. Krombein (1$, 
USNM). 
Description 
Male. Body length 5.5 mm. Anepistemal disc present. First 
flagellomere orbicular. Bristles black. Arista pubescent. 
Vibrissa relatively long and curved. Mid fronto-orbital 
absent. Ocellar bristle minute. Two dorsocentral bristles with 
small bristle in front of anterior dorsocentral. Gena shallow 
and not sharply angled. Face and buccal cavity pilose and 
meeting at an angle. Head dark brown with apex of palpus 
and antenna (excluding arista) yellow, with anterior margin of 
first flagellomere infuscated; gena with thin dorsal tomentose 
strip; frons shiny. Thorax dark brown. Legs yellow with base 
of fore coxa brown, fore tarsi and tibia dark brown, apex of 
fore femur brown, apex of hind femur brownish and base of 
hind tibia brown. Fore tarsi strongly compressed laterally. 
Abdomen dark brown. Wing dark along anterodistal margin 
and between R 4+5 and M, to level of dm-cu; lightly clouded 
around CuAj. M 1+2 ratio 6.0. Halter white with base and side 
of stalk infuscated. Paratype male differs as follows: face, 
parafacial and upper half of gena dark yellow; fore coxa 
entirely light yellow; fore femur yellow with faint inner-distal 
spot; hind legs entirely yellow; wing dusky, except along 
posterior and anterobasal margins. Halter brown. 
Female. As described for Australian male except antenna 
entirely dark brown and pigment on hind leg slightly darker. 
Male terminalia (Figs 28-31). Epandrium as wide as high 
and length % height. Cerci entirely fused, large and ovate. 
Surstylus length approximately half height of epandrium, 
thin, pointed and relatively anterior in position; several 
tubercle-like bristles on inner face apically. Phallapodeme 
nearly as long as hypandrium. Hypandrium + pregonite 
with several distal and basal setulae, and with triangular 
anterior and lobate basal lobes. Ribs of phallus nearly equal 
in length, both becoming wider and convoluted apically; one 
rib S-shaped apically and with medial projection produced 
into two overlapping triangular points. 
Female terminalia (Fig. 92). Spermathecae spherical, 
not strongly sclerotized and without apical invagination. 
Remainder of internal terminalia not visible. 
Etymology. The specific name refers to the presence of only 
two fronto-orbital bristles in this species. 
Comments. The presence of only two pairs of fronto-orbital 
bristles is diagnostic of this species, as is the pattern on the 
wing, the ovate male cerci and the crossing spines on the 
phallus. Heteromeringia bisetosa is the only Australian 
species of that genus known to extend outside of Tasmania 
and continental Australia. 
Heteromeringia digitula n.sp. 
Figs 13, 41-43 
Type material. Holotype Queensland: 12.44S 143.14E, 3km ENE Mt. 
Tozer, 28.vi-4.vii. 1986, D.H. Colless (1(3, ANIC). Paratypes Queensland: 
12.44S 143.14E, 3km ENE Mt. Tozer, 28.vi^l.vii.l986, D.H. Colless (1?, 
ANIC), Malaise trap, ethanol (1^, DEBU). 
Description 
Male. Body length 3.3-4.3 mm. Anepistemal disc present. 
First flagellomere orbicular. Bristles brown. Arista pubescent. 
Vibrissa relatively long and curved. Ocellar bristle minute. 
Two dorsocentral bristles. Gena shallow and not sharply 
angled. Head yellow with first flagellomere infuscated on 
distal half of inner face, ocellar tubercle brown, gena with 
small medial msty spot, anterior half of occiput and upper 
half of gena light yellow and silvery tomentose, upper- 
posterior half of occiput dark brown, back of head with one 
pair of dorsolateral spots and base of palpus whitish; frons 
tomentose. Thorax yellow with two very thin postsutural 
stripes (along dorsocentral rows, continuing onto scutellum), 
faint posterolateral spots, orange tint in posterolateral comer 
of scutum, thin sub-scutellar stripe on laterotergites and 
orange mottling on pleuron. Fegs yellow with fore tibia 
and tarsi dark brown (knee yellow), coxae light yellow to 
white, and fore femur with oblique dark brown inner-distal 
spot. Fore tarsi strongly compressed laterally. Abdomen dark 
brown with segments 1 and 2 yellow. Wing dusky along 
anterodistal margin at R 2+3 and R 4+5 . Halter white. 
Female. As described for male except as follows: first 
flagellomere entirely yellow; spots on back of head only 
present near margins; tergites 1-4 yellow with dark medial 
stripe and tergite 4 also with posterolateral spot; tergite 8 
(infuscated anterodorsally) and terminalia yellow; wing 
clear around R 4+5 and lightly clouded around Mj past r-m. 
Male terminalia (Figs 41-43). Epandrium width nearly 
twice length and height. Sides of cerci nearly parallel; with 
deep medial emargination. Surstylus subquadrate with length 
and height approximately 2 h that of epandrium; inner-distal 
margin lined with tubercle-like bristles. Phallapodeme nearly 
as long as hypandrium. Hypandrium+pregonite divided 
into thin anterior lobe with one stout bristle and wider 
posterior lobe with numerous setulae. Basiphallus with thin 
posterobasal lobe. Ribs of phallus nearly equal in length 
with one rib bifid apically and one rib pointed and sharply 
recurved medially. 
Female terminalia. Similar to those of limacens (Fig. 100). 
Etymology. The specific name refers to the recurved finger¬ 
like process on the phallus. 
Comments. See comments for Heteromeringia hypoleuca. 
