Lonsdale: Australian Heteromeringia flies (Clusiidae) 
243 
Heteromeringia hardyi D.K. McAlpine, 1960 
Figs 1,2, 6, 10, 32-34, 93 
Heteromeringia hardyi D.K. McAlpine, 1960: 75. 
Type material. Holotype: New South Wales: Blue Mountains, Katoomba, 
19.xii.1956, G.H. Hardy (1(3, AMS). Paratypes: New South Wales: Same 
collection as holotype (1$, AMS), 6.xii.l956 (1$, AMS), 29.xii.1956 (13, 
AMS), 23.xi. 1956 (1$, AMS), 17.xii. 1956 (1 $, AMS). 
Additional material examined. New South Wales: Katoomba, 24.i. 1961, 
G.H. Hardy (13, AMS), Tahmoor, 7.X.1978, B.J. Day (1$, AMS), 
20.ix.1981, D.K. McAlpine & B.J. Day (1$, AMS), Mt. Banks, Blue 
Mountains, l.i.1982, D.K. McAlpine (1$, AMS), 8km S Mt. Wilson, 
Blue Mts., 27.ix. 1989, D.K. McAlpine (3$$, AMS), 3km E Wedderbum, 
34°08'S 150°49’E, 28.X.2003, at MV light, D. Britton (1$, AMS), Mooney 
Mooney Ck. Near Gosford, 17.xii. 1981, B. Day (1<3, AMS), Werrikimbe 
NP, 31°11'24"S 152°09'39"E, 1030 m, E. Tasker, l-7.xii.1997, sticky trap 
on E. viminalis, WC-WT-127-6 (1$, AMS), Werrikimbe NP, 31°11'56"S 
152°10'23"E, 1025 m, E. Tasker, 1-7.xii. 1997, sticky trap on E. cameronii, 
WC-WN-127-3 (1$, USNM), Carrai SF, 30°59’45"S 152°16’23"E, 930 
m, E. Tasker, 3-8.xii.1997, sticky trap on E, campanulata, CS-FZ-127-6 
(13, AMS), Carrai SF, 30°54’35"S 152°16’26"E, 1090 m, E. Tasker, 3-8. 
xii. 1997, sticky trap on E. campanulata, CC-FK-127-4 (1(3, AMS), Carrai 
SF, 30°54'35"S 152°16'26"E, 1090 m,E. Tasker, 11-16.L 1998, sticky trap on 
E. obliqua, CC-FK-018-2 (13, AMS), Carrai SF, 30°54’33"S 152°16’28"E, 
1075 m, E. Tasker, 3-8.xii.1997, sticky trap on E. campanulata, CC-CR- 
127-2 (13, AMS), Carrai SF, 30°54T9"S 152°17'36"E, 1055 m, E. Tasker, 
3-8.xii. 1997, sticky trap on E. campanulata, CC-DP-127-4 (13 1 $, AMS), 
Carrai SF, 30°54T9"S 152°17’36''E, 1055 m,E. Tasker, 3-8.xii.1997, sticky 
trap on E. obliqua, CC-DP-127-1 (13 1$, USNM), Carrai SF, 30°54'33"S 
152°16'28''E, 1075 m, E. Tasker 11-164.1998, sticky trap on is. cameronii, 
CS-FZ-127-4 (1$, AMS), Carrai SF, 30°54T9"S 152°17'36"E, 1055 m, 
E. Tasker, 3-8.xii.1997, sticky trap on E. cameronii, CC-DP-127-6 (13 
1$, USNM), Carrai SF, 30°54'19"S 152°17'36"E, 1055 m, E. Tasker, 
11-164.1998, sticky trap on E. obliqua, CC-DP-128-4 (1$, AMS), Carrai 
SF, 30°54T9"S 152°17'36"E, 1055 m, E. Tasker, 11-164.1998, sticky trap 
on E. obliqua, CC-DP-018-4 (13, DEBU). Queensland: Mt. Glorious, 
27°19'54"S 152°45'29"E, Malaise, T. Hiller, 24-30.xi.1997 (1$, DEBU), 
3-94.1998 (1$, DEBU), Mt. Glorious, l.ix-17.x,1990, A. Hiller, Malaise 
(1$, ANIC), N QLD, 2 mi W Paluma, 3000', 164.1970, MV lamp, G.A. 
Holloway (1$, AMS). 
Redescription 
Male (Figs 1, 2, 6). Body length 3.0-4.5 mm. Anepisternal 
disc present. First flagellomere recessed dorsally and lobate 
ventrally. Bristles dark brown. Arista pubescent. Vibrissa 
short and straight. Ocellar bristle relatively stout and longer 
than tubercle. Two dorsocentral bristles. Gena high and 
sharply incurved medially at mid-height. Face and buccal 
cavity evenly curved and velvety. Frons brown on posterior 
% (lateral and posterior margins paler); back of head dark 
brown; occiput and posteroventral comer of gena dark brown; 
face lighter dorsally; gena silvery tomentose above genal 
bristles; anterior % of frons pilose. Thorax dark brown. Legs 
yellow except as follows: mid coxa brown dorsally; hind 
coxa light brown; fore femur with large, dark anterodistal 
and posterodorsal spots (usually somewhat confluent); fore 
tarsi brownish on distal two or three segments; hind femur 
somewhat dusky on outer surface apically; hind tibia brown 
on basal Vs; mid and hind femora sometimes brownish 
dorsomedially; mid femur and mid and hind tibiae sometimes 
with wide central band; fore coxa sometimes with minute 
brownish spot on dorsal margin, and gena and venter of face 
dusky. Fore tarsi cylindrical in cross section. Abdomen dark 
brown. M 1+2 ratio 6.5-13. Wing clouded on distal half of 
anterior margin; veins, excluding costa and R 2+3 , surrounded 
by light infuscation. Halter white with base and side of stalk 
infuscated. 
Female. As described for male except as follows: fore tarsi 
dark brown, high and laterally flattened (height sharply 
decreasing past second segment); fore femur with outer-distal 
spot absent or fused to inner spot; fore tarsi dark brown; 
mid coxa brown on outer-dorsal half; fore tibia sometimes 
white with base dark brown; mid tibia sometimes mottled 
medially; apex of mid and hind femora and base of mid tibia 
variably infuscated; base of hind tibia brown; frons pilose 
anteromedially; first flagellomere dark along anterior and 
dorsal margins, or entirely dark excluding basal margin; gena 
and venter of face dusky; tergites 2-5 sometimes yellow on 
anterior Vs, or with anterior half of tergite 2 and all of tergites 
3-5 yellow; wing without light infuscation around veins. 
Two non-type females with fore tarsi only slightly modified, 
fore tibia entirely light yellow, apex of fore femur brown, 
hind knee brown and face and gena entirely reddish-brown. 
Male terminalia (Figs 32-34). Epandrium deeply excavated 
on anterior margin, continuous with posterior margin of 
large hypandrium+pregonite. Cerci widest subapically 
and emarginate. Surstylus long and thin (narrowest 
subapically) with few anterodistal setulae and several inner- 
distal tubercles. Hypandrium+pregonite with single large 
basal lobe with anterodistal and medial setulae, two stout 
medial bristles and stout posterodistal bristle on inner face. 
Distiphallus with one rib short, thin and tapered, and one rib 
large and elaborate apically: with thin posteriorly-directed 
process, one wider anteriorly-directed process and elongate 
bifid distal process. 
Female terminalia (Fig. 93). Spermatheca with several 
shallow basal wrinkles and many minute medial tubercles; 
apex broadly rounded with apical invagination small and 
offset. Ventral receptacle and flagellum not visible. 
Heteromeringia helina n.sp. 
Figs 35-37, 94 
Type material. Holotype: New South Wales: Carrai SF, 30°54'33"S 
152°16'28"E, 1075 m, E. Tasker, 3-8.xii. 1997, sticky trap on E. 
campanulata, CC-CR-127-2 (13, AMS). Paratypes: New South Wales: 
Carrai SF, 30°54T9"S 152°17'36"E, 1055 m, E. Tasker, 3-8.xii.1997, 
sticky trap on E. campanulata, CC-DP-127-4 (233, AMS), Werrikimbe 
NP, 31°11'56"S 152°10'23''E, 1025 m, E. Tasker, l-7.xii.1997, sticky 
trap on E. viminalis, WC-WN-127-5 (13, DEBU), Werrikimbe NP, 
31 0 H'56''S 152°10'23"E, 1025 m, E. Tasker, l-7.xii.1997, sticky trap on 
E. campanulata, WC-WN-127-1 (3, AMS; 13, USNM), sticky trap on E. 
saligna, WC-WN-127-6 (13, AMS). 
Additional material examined. New South Wales: Leura Falls, 34.1973, 
D.H. Colless (2$?, ANIC). 
Description 
Male. Body length 2.9-3.4 mm. Anepisternal disc present. 
First flagellomere orbicular. Bristles black. Arista pub¬ 
escent. Vibrissa relatively long and curved. Ocellar bristle 
long and as thick as postvertical. Two dorsocentral bristles. 
Gena relatively small and angled inwards, but not bent at 
mid-height. Face and buccal cavity smoothly curving and 
velvety. Frons brownish-orange with anterior, lateral and 
posterior margins yellowish; first flagellomere light yellow; 
