248 
Records of the Australian Museum (2009) Vol. 61 
Heteromeringia laticornis D.K. McAlpine, 1960 
Figs 14, 47-49, 99 
Heteromeringia laticornis D.K. McAlpine, 1960: 76. 
Type material. Holotype: New South Wales: National Park, 19.iii. 1957, 
D. K. McAlpine (13, AMS). Paratype: New South Wales: Otford, 
12.X.1957, D.K. McAlpine (1$, AMS). 
Additional material examined. New South Wales: E NSW, Royal national 
Park, E Waterfall rail.stat., 12.X.2002,34.09S 151.01E, B. Merz, diff. types 
forest (233, MHNG), Otford, 18.L1964, D.K. McAlpine (1 3, AMS), 
Mooney/Mooney Creek near Gosford, 29.ii.1975, D.K. McAlpine (13, 
AMS), Tubrabucca, Upper Hunter Dist., 21.xii.1975, G. Daniels (1$, AMS), 
“Lorien”, 3km N Lansdowne, rainfor. edge, Malaise, 15—22.ii. 1987, G. 
Williams (1^, AMS), Carrai SF, 30°59’45"S 152°16’23 ,r E, 930 m, E. Tasker, 
3-8.xii.1997, sticky trap on E. cameroni, CS-FZ-018-2 (333 1$, AMS), 
Carrai SF, 30°54’19"S 152 0 17'36"E, 1055 m, E. Tasker, 11-16.1.1998, 
sticky trap on E. campanula, CC-DP-018-2 (333, AMS), Carrai SF, 
30°54'19"S 152°17'36"E, 1055 m, E. Tasker, 11-164.1998, sticky trap on 
E. obliqua, CC-DP-018-4 (1$, AMS; 1$, USNM), Carrai SF, 30°54’35"S 
152 0 16'26"E, 1090 m, E. Tasker, 11-164.1998, sticky trap on E. obliqua, 
CC-FK-018-5 (233, AMS), Carrai SF, 30°54’35"S 152°16’26"E, 1090 m, 
E. Tasker, 11-164.1998, sticky trap on E. campanulata, CC-FK-127-4 (1 V 
AMS), Carrai SF, 31°00T9"S 152°16’24"E, 940 m, E. Tasker, 3-8.xii.1997, 
sticky trap on E. campanulata, CS-GP-018-4 (333 1$, AMS), Carrai SF, 
30°59'45"S 152°16’23"E, 930 m, E. Tasker, 3-8.xii.1997, sticky trap on 
E. campanulata, CS-FZ-127-3 (133 2$$, AMS), Carrai SF, 30°54T9"S 
152 0 17'36"E, 1055 m, E. Tasker, 3-8.xii. 1997, sticky trap on E. obliqua, 
CC-DP-127-5 (1$ 13, DEBU), Carrai SF, 30°58'48"S 152°17'06"E, 975 m, 
E. Tasker, 3-8.xii.1997, sticky trap onE. viminalia, CS-RO-127-2 (1 3 1?, 
AMS), Carrai SF, 30°58’48"S 152°17’06"E, 975 m, E. Tasker, 11-164.1998, 
sticky trap on E. obliqua, CS-RO-018-5 (233, AMS; 1$, DEBU), Carrai 
SF, 30°54'33''S 152°16’28"E, 1075 m, E. Tasker, 11-164.1998, sticky trap 
on E. campanulata, CC-CR-018-6 (2 33, AMS; 1 3, DEBU), Carrai SF, 
30°54'33"S 152°16'28"E, 1075 m, E. Tasker, 3-8.xii.1997, sticky trap 
on E. campanulata, CC-CR-127-2 (13, AMS), Carrai SF, 30°54'19"S 
152°17’36"E, 1075 m, E. Tasker, 3-8.xii.1997, sticky trap on E. cameronii, 
CC-DP-127-6 (13, USNM), Carrai SF, 30°54'19"S 152°17’36''E, 1055 
m, E. Tasker, 3-8.xii.1997, sticky trap on E. campanulata, CC-DP-127-4 
(233 2$.§VAMS), Werrikimbe NP, 31°12'00"S 152°09'00"E, 1060 m, E. 
Tasker, l-7.xii.1997, sticky trap on E. obliqua, WC-MR-127-2 (1^, AMS), 
Werrikimbe NP, 31° 16’50"S 152°03T9"E, 1045 m,E. Tasker, l-7.xii.1997, 
sticky trap on E. campanulata, WS-FC-127-1 (233, AMS), Werrikimbe 
NP, 3ri6'50"S 152°03'19"E, 1045 m, E. Tasker, l-7.xii.1997, sticky trap 
onE. saligna, WS-FC-127-5 (lc^, AMS). Northern Territory: East Jabiru, 
24-28.v. 1988, A. Wells, yellow pan trap (1$, ANIC). Queensland: trib. 
Krombit Ck., Krombit Tops SF, gully r.f. & wet sclera., 31.x-2.xi.1997, 
sticky trap on E. saligna, D. Bickel (1 AMS). South Australia: Mt. 
Barker (summit), iii-iv. 1988, A.D. Dustin, M/trap (4$ ANIC). Tasmania: 
Rosebery, ii. 1977, BPBM (1$, AMS). 
Redescription 
Male. Body length 2.4-3.0 mm. Anepisternal disc present. 
First flagellomere recessed dorsally and lobate ventrally (as 
in Figs 3,4). Bristles black. Arista pubescent. Vibrissa short 
and straight. Ocellar bristle long and thin. Two dorsocentral 
bristles. Gena high and sharply incurved medially at mid¬ 
height. Face and buccal cavity evenly curving and velvety. 
Head yellow with posterior Vs of frons (excluding yellowish 
lateral and posterior margins), back of head, occiput, clypeus 
and posterior Vs of gena dark brown; first flagellomere 
variable, ranging from entirely yellow to entirely dark brown 
(excluding base); dorsal half of gena silvery tomentose; 
anterior Vs of frons pilose. Thorax dark brown. Fore femur 
dark brown on distal Vs (slightly more extensive on inner 
face); hind femur dark brown on distal half (excluding knee); 
fore tarsi dark brown, sometimes with distal segment lighter; 
fore tibia lighter on basal half with inner apex brownish; fore 
coxa white with dark yellow to light brown spot on outer 
base (sometimes developed into elongate brown stripe or 
fore coxa brown on posterior half); mid coxa dark brown on 
outer-dorsal half; remainder of legs yellow; sometimes legs 
(excluding fore coxa) dark brown with base of femora, knees 
and apex of mid and hind tibiae yellow, and mid and hind tarsi 
light brown. Fore tarsi ovate in cross section. Abdomen dark 
brown. M 1+2 ratio 7.3. Wing with anterodistal infuscation; 
remainder of wing sometimes dusky. Halter white with base 
and side of stalk infuscated. 
Female. As described for male except as follows: first 
flagellomere slightly more orbicular; mid and hind femora 
dark brown with base and apex yellow; fore tibia entirely 
dark brown excluding yellow base; mid and hind tibiae 
sometimes dark brown excluding base and apex, or entirely 
yellow; mid and hind tarsi light brown to yellow; fore tarsi 
with slight to pronounced lateral compression; frons shiny 
and black with anterior margin yellow (sometimes more 
extensively orange or yellow on anterior and anterolateral 
margins); palpus occasionally brownish apically. 
Male terminalia (Figs 47-49). Height and length of 
epandrium Vs width. Cerci relatively small and rounded 
with shallow apical emargination. Surstylus small and 
rounded with numerous inner-distal tubercle-like bristles. 
Hypandrium+pregonite with large anterior lobe with two 
stout bristles, and small posterior lobe with several setulae. 
Phallus with thin longitudinal sclerite at midpoint, stouter 
laterally-projecting subapical sclerite (width varying), and 
large U-shaped apical sclerite with two basal processes. 
Female terminalia (Fig. 99). Tergite 8 wrapping around 
segment posteriorly, enclosing sternite 8 distally. Ventral 
receptacle narrowing apically and flagellum very long and 
straight. Spermatheca as wide as long, smooth and with 
several basal wrinkles. 
Comments. The species Heteromeringia laticornis, H. 
hardyi and H. magnicauda appear to be closely related on the 
basis of a number of putative synapomorphies: the vibrissae 
are short and straight, the gena is relatively high and sharply 
curved inwards, and the first flagellomere is lobate ventrally 
and recessed dorsally (sometimes indistinct), shifting the 
arista closer to the pedicel. Heteromeringia helina and 
H. trisetosa may be basal to this group since they share a 
similar gena and the male fore tarsi are similarly ovate in 
cross section; the distiphallus and hypandrium of H. helina 
are also highly similar to those of H. hardyi. 
