Lonsdale: Australian Heteromeringia flies (Clusiidae) 
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and lateral and posterior margins yellowish; first flagellomere 
with infuscation at base of arista; occiput, back of head and 
posteroventral corner of gena dark brown; dorsal half of 
face light yellow; buccal cavity, anterior portion of frons and 
dorsal Vs of gena pilose; mentum sometimes dark brown. 
Thorax dark brown. Fore tibia dark brown on distal Vs and 
fore femur light brown (darker on basal Va-Vi) with knee 
yellow; fore coxa brownish at base; fore tarsi (ovate in cross 
section) yellow with distal two tarsomeres light brown (QLD 
specimens with all segments brown); dorsal margin of mid 
coxa brown to orange; mid and hind femora brown medially, 
mid tibia brown medially on inner face and hind tibia brown 
on basal half (sometimes mid and hind femora and tibiae 
entirely brown excluding base and apex); hind coxa brown; 
remainder of legs yellow. Abdomen dark brown. M 1+2 ratio 
6.4. Wing with anterodistal infuscation. Halter white with 
base and side of stalk infuscated. 
Female. Unknown. 
Male terminalia (Figs 54-57). Epandrium bulbous and 
larger than pregenitalic segments. Cerci rounded laterally 
and with apex narrow and emarginate. Surstylus triangular 
and strongly narrowed apically; bare on outer face and with 
inner-distal tubercle-like bristles. Phallapodeme relatively 
long and stout. Hypandrium+pregonite setulose medially 
and distally with two stout anteromedial bristles (inner-distal 
face with additional stout bristle); flat along anterior face and 
with suture on distal Vs; distal margin notched. Basiphallus 
with dorsal process at point of attachment to distiphallus 
and fused to hypandrium+pregonite posterolaterally. Ribs 
of distiphallus of equal length, fused medially and with one 
pair of small membranous wings. 
Etymology. The specific name refers to the strikingly large 
(Latin magnus ) male genitalia (Latin cauda ) of this species. 
Comments. Although the male genitalia of this species 
are incredibly diagnostic—particularly the bulbous male 
epandrium—the only other potentially autapomorphic 
character is relatively pale fore tarsi. The fore tarsi are 
brown in the male from Queensland, however, and they 
are potentially brown in the as yet undiscovered females, 
since female Heteromeringia are often darker than their 
respective males, particularly on the fore legs. This suggests 
the possibility that females of H. magnicauda are inseparable 
from those of the similar H. laticornis. 
Heteromeringia montana n.sp. 
Figs 58-61 
Type material. Holotype: Queensland: Kuranda, 6.X.2002, A. Freidberg 
(1$, AMS). Paratype: Queensland: 12.43S 143.18E, llkmEnEofMt. 
Tozer, ll-16.vii.1986, Malaise trap, D.H. Colless (Id 1 , ANIC). 
Additional material examined [“species A” of D.K. McAlpine (I960)]: 
New South Wales: Bronte, near Sydney, 31.iii.1958 (Id, AMS), Otford, 
Illawarra District, 26.U959 (1$, AMS). 
Description 
Male. Body length 2.1 (NSW male), 3.0 (female), 3.5 mm 
(QLD males). Anepisternal disc present. First flagellomere 
orbicular. Bristles black. Arista closely pubescent. Vibrissa 
relatively long and curved. Ocellar bristle thin and longer 
than tubercle. Two dorsocentral bristles with small bristle 
in front of anterior dorsocentral. Gena relatively high and 
bent medially. Face and buccal cavity pilose and meeting 
at an angle. Frons dark brown medially, fading to orange 
anteriorly, and with lateral margins yellowish; back of head 
dark brown above foramen; occiput dark brown (excluding 
venter); face brownish centrally; first flagellomere brown on 
ventral 3 A of anterior margin and on ventral half of inner face; 
remainder of head yellow; gena pilose on dorsal Vs; anterior 
half of frons tomentose. Thorax dark brown. Legs yellow 
with mid and fore coxae dark brown dorsally, fore tibia 
brownish (darker apically), fore tarsi brown (and laterally 
compressed), fore femur with inner-distal spot and mid and 
hind femora with light, sparse mottling on posterior surface 
(sometimes forming complete band). Abdomen dark brown. 
M 1+2 ratio 5.5-6.0. Wing with anterodistal infuscation. Halter 
white with colour of base and stalk indiscernible. NSW male 
differs in that distal Vs of femur brown, back of head and 
occiput entirely brown, inner-distal face of first flagellomere 
lightly infuscated, and face yellow. 
Female. As described for type males except as follows: fore 
tibia evenly brown; fore femur with light outer-distal spot; 
mid femur and coxa entirely yellow; frons shiny and brown 
on posterior Vs; first flagellomere infuscated on distal half 
of inner face dorsally. 
Male terminalia (Figs 58-61). Epandrium wider than long 
or high. Cerci widest medially and emarginate. Surstylus 
very dark and well sclerotized; elongate triangular in 
outline with apex rounded and curved inwards; anterior 
margin bare; tubercle-like bristles on inner-distal margin. 
Hypandrium+pregonite divided longitudinally by thin suture 
(dorsal section with several posterior setulae and ventral 
section recessed and bifid with several posterodistal setulae 
and two stout anterior bristles). Phallapodeme as high as 
hypandrium+pregonite and clavate (seen laterally). One rib 
of distiphallus ending medially; second rib with complex 
distal branching. 
Female terminalia. Not dissected. 
Etymology. The specific name is Latin for “mountain”, and 
refers to the collection locality of the ANIC type specimen. 
Comments. While the identity of the two specimens 
designated “species A” by D.K. McAlpine (1960) is still 
uncertain, they appear to be most closely allied to the 
above males examined from Queensland, and are here 
tentatively included in Heteromeringia montana. The male 
(not dissected) is similar to the types in that the fore tibia 
is entirely brown, the fore tarsi are ovate in cross section, 
the mid and hind femora are entirely yellow, the vibrissa is 
relatively long and the surstylus is more tapered apically. 
This male, however, is relatively small (2.1 mm), the gena 
is similar to that seen in H. laticornis , the first flagellomere 
is dorsally (not ventrally) infuscated, and the frons and 
fore femur are darker. The female is much more similar in 
appearance to the type specimens (particularly with regards 
to the colour of the head and legs), but because of the 
difficulty of associating Australian Heteromeringia females 
with males, this slightly dissimilar specimen is not included 
in the type series. 
