Lonsdale: Australian Heteromeringia flies (Clusiidae) 
253 
The genitalia are most similar to those of Heteromeringia 
macropa, but the surstylus is longer and more gradually 
tapered with additional tubercle-like bristles, the hypandrium 
differs in shape and chaetotaxy, and the phallus differs in 
the number and shape of medial processes. The surstylus 
is also similar to that of H. helina, but the internal genitalia 
are quite different. Externally, the type specimens can be 
mistaken for H. patula, but the anepisternum is shiny (not 
tomentose), the fore tibia is entirely brown, the frons and 
first flagellomere are darker and the scutum is not pale in 
the posterolateral corner. 
Heteromeringia norrisi D.K. McAlpine, 1960 
Figs 62-64 
Heteromeringia norrisi D.K. McAlpine, 1960: 74. 
Type material. Holotype: Western Australia: Applecross, 10.vi.1934, 
K.R. Norris, emerged from rotting wood 18.vii.1934 (Id', ANIC). 
Redescription 
Male. Body length 3.0-3.8 mm. Anepisternal disc present. 
First flagellomere orbicular. Bristles black. Arista closely 
pubescent. Vibrissa curved and relatively short (although 
not as short as vibrissa of H. laticornis). Ocellar bristle long. 
Two dorsocentral bristles (broken) with slightly more well 
developed setula in front of anterior dorsocentral. Occiput 
relatively large, and gena high and gradually curved medially. 
Face and buccal cavity evenly curving and velvety. Frons 
dark brown with anterior margin, posterior margin and 
faded spot in front of anterior ocellus brownish-orange; 
occiput and posterior half of gena brown; antenna (excluding 
arista) orange-yellow with anterodorsal margin and distal Vs 
of first flagellomere infuscated; distal Vs of palpus brown, 
(excluding inner margin); parafacial and dorsal half of gena 
silvery tomentose, frons broadly pilose anteriorly. Thorax 
dark brown. Fore legs dark brown with base of tibia yellow 
and fore coxa dirty yellow/light brown and silvery tomentose 
with base brown; mid tibia, coxa and femur dark brown with 
base and apex of femur yellow; mid tarsi yellow; hind coxa 
brown (remainder of hind legs missing—D.K. McAlpine 
[1960] describes those parts as dark brown with knee and 
middle of tarsi yellowish). Fore tarsi ovate in cross section. 
Abdomen dark brown. M 1+2 ratio approximately 5.5. Wing 
only clouded along R 2+3 and R 4+5 on distal Vs. Halter white 
with stalk brownish (darker to base). 
Female. Not examined. 
Male terminalia (Fig. 64). Epandrium largely obscured, 
but width, height and length appearing subequal. Cerci 
approximately half height of epandrium, narrowing distally 
and with small apical emargination. Surstylus long and 
triangular (strongly tapered on distal half) with several distal 
tubercle-like bristles. 
Comments. Heteromeringia norrisi is one of only two 
Heteromeringia known from Western Australia. The other 
is the relatively widespread H. trisetosa, which also occurs 
in New South Wales, Queensland and Tasmania, and is 
characterized by three pairs of dorsocentral bristles. 
One of the paratype females (not examined) was collected 
in a log with termites (D.K. McAlpine, 1960). 
Heteromeringia patula n.sp. 
Figs 65-68, 102 
Type material. Holotype: New South Wales: Royal Natl. Pk., Scientists’ 
Cabin Trail, creek bed, sweep, 25.xii.2003, S.A. Marshall (Id', AMS). 
Paratypes: New South Wales: same collection as holotype (Id, DEBU), 
Otford, NSW, 29.ii. 1964, D.K. McAlpine (Id, AMS). 
Additional material examined. New South Wales: Carrai SF, 
30°54’19"S 152°17’36"E, 1055 m, E. Tasker, 3-8.xii.1997, sticky trap on 
E. campanulata, CC-DP-127-4 (2$?, AMS), Werrikimbe NP, 31°11'56"S 
152°10'23"E, 1025 m, E. Tasker, l-7.xii.1997, sticky trap on E. saligna, 
WC-WN-127-6 (1 $, AMS). 
Description 
Male. Body length 3.2-3.4 mm (male) and 4.0-5.8 mm 
(female). Anepisternal disc present. First flagellomere 
orbicular. Bristles black. Arista closely pubescent. Vibrissa 
relatively long and curved. Ocellar bristle thin and longer 
than tubercle. Two dorsocentral bristles with small bristle in 
front of anterior dorsocentral. Gena shallow and not strongly 
angled or bent. Face and buccal cavity pilose and meeting 
at an angle. Posterior half of frons dark brown with lateral 
and posterior margins paler, and anterior half tomentose and 
bright yellow to orange anteriorly and laterally; occiput and 
back of head above foramen dark brown; distal margin and 
ventral Vi- 3 A of inner face of first flagellomere infuscated; 
ventral margin of gena shiny orange; remainder of head 
yellow; dorsal Vs of gena pilose. Thorax dark brown. Fore 
coxa white with outer base yellow to brown; mid coxa brown 
laterally; fore tarsi dark brown and ovate in cross section; 
fore tibia mottled brown laterally; fore femur with dark inner- 
distal spot and with variable brown mottling along length; 
basal Vs of mid and hind femora yellow, mottled brown, 
or with faint posterodorsal spot; remainder of legs yellow. 
Abdomen brown with tergite 1 yellow. M 1+2 ratio 4.0-8.8. 
Wing with light anterodistal infuscation. Halter white with 
base and side of stalk infuscated. 
Female. As described for male except as follows: 
postpronotum and notopleuron paler; mid coxa light brown 
dorsally; fore tibia dark brown; fore femur light yellow on 
basal half and dark brown on distal half; mid femur with light 
brown medial mottling; hind femur with nearly indistinct 
medial mottling; frons brownish orange (yellow laterally); 
apex of palpus infuscated; first flagellomere yellow with 
anterior and inner-distal margin brown; occiput yellow; gena 
and face bronze and frons yellow with medial bronze/brown 
tint; abdomen entirely brown; wing infuscated along distal 
half of anterior margin. 
Male terminalia (Figs 65-68). Epandrium slightly higher 
than long and distinctly wider than high. Surstylus short and 
broadly triangular with apex rounded and posterior comer 
long and fused to epandrium. Cerci tapering apically and 
shallowly emarginate. 
Hypandrium+pregonite with two stout bristles on anterior 
lobe, and very dark elongate posterior sclerite (wide and 
truncate distally with several posteromedial setae and several 
inner-distal setulae) that strongly projects laterally and is 
distinct even in non-dissected specimens. Phallapodeme 
thin, but relatively long, arched and broad distally. Ribs of 
distiphallus long, parallel and twisted medially. 
