260 
Records of the Australian Museum (2009) Vol. 61 
Heteromeringia stegna n.sp. 
Figs 81-84 
Type material. Holotype: Northern Territory: Litchfield Natl. Pk., 
20.L1998, A. Zwick (1(3, AMS). Paratype: Queensland: Bamaga-Capt 
Billy CkRdjnct 16kmNEHeathlandsH.S„ 11°41'S 142°42'E, 16.iii.1992, 
G. Daniels & M.A. Schneider (1 S, UQIC). 
Description 
Male. Body length 3.1 mm. Anepisternal disc present. 
First flagellomere orbicular. Bristles black. Arista closely 
pubescent. Vibrissa relatively long and curved. Ocellar 
bristle minute. Two dorsocentral bristles with small bristle 
in front of anterior dorsocentral. Gena shallow and not 
strongly angled or bent. Face and buccal cavity pilose and 
meeting at an angle. Frons black and shiny with posterior 
margin yellowish, and lateral margin yellow and pilose (or 
pilose anteromedially); back of head, occiput and posterior 
2 /3 of gena dark brown; clypeus and mentum brown; anterior 
margin of first flagellomere and distal half of inner face 
brown; basal 2 /3 of palpus brown (holotype) or entirely 
dark brown excluding apex; remainder of head yellow 
(holotype) or orange-brown; dorsal margin of gena pilose. 
Thorax dark brown. Legs yellow with fore tarsi and tibia 
dark brown; fore femur dark brown on distal Vd (holotype) 
or with inner-distal spot. Fore tarsi very slightly compressed 
laterally. Abdomen brown. M 1+2 ratio 5.3. Wing with small 
anterodistal infuscation (holotype), or dusky with base and 
posterior margin clear. Knob of halter brown, and base and 
side of stalk infuscated. 
Female. Unknown. 
Male terminalia (Figs 81-84). Epandrium elongate oval 
with margins nearly meeting distally. Surstylus subtriangular, 
small, setulose on inner face and with base fused to anterior 
margin of epandrium. Cerci arched over surstyli, setose, 
widest at apex, and shallowly but broadly emarginate. 
Hypandrium+pregonite bilobed with broad anterior lobe 
with two stout bristles, and linear posterior lobe with several 
posterodistal setulae. Phallapodeme as long a hypandrium. 
Distiphallus with strong medial curve where one rib ends 
and the other rib extending into a long process accompanied 
by irregular U-shaped sclerite. 
Etymology. The specific name is Latin for “constricted”, in 
reference to the apical narrowing of the epandrium. 
Comments. While relatively non-descript externally, the 
male genitalia of this species are heavily modified: the 
epandrium is constricted apically, the cerci are very broad 
apically, and the nearly vestigial surstylus is positioned 
anteriorly (not apically) and partially fused to the epandrium. 
Heteromeringia trisetosa n.sp. 
Figs 9, 85-87, 106 
Type material. Holotype: Tasmania: 41.50S 146.03E, Pelion Hut 
3km S Mt. Oakleigh, 850 m, 8.i-12.ii.l991, A. Calder & W. Dressier, 
Malaise #5, closed forest (1<3, ANIC). Paratypes: New South Wales: 
Penrose SF, nr. Marulan, 20.x. 1989, D.K. McAlpine (1(3, AMS), Mooney 
Mooney Creek near Gosford, 263.1984, B. Duckworth, B. Day and D.K. 
McAlpine (1(3, AMS), E NSW, Royal National Park, E Waterfall rail. 
Stat., 12.X.2002, 10, 34.09S/151.01E, B. Merz/ 10, diff. types forest (1?, 
MHNG). Queensland: SE QLD, Brisbane, Griffith Uni., Nathan Campus, 
23.X.2002, 18, 27.33S/153.04E, dry & wet sclerophyll forest, ferns, B. 
Merz / 18 (1(3, MHNG). Western Australia: Shannon townsite camp, 
shrimp carrion trap, 25.xi.2003, S.A. Marshall (1(3, DEBU). Tasmania: 
Pelion Hut, 30.xi.l990-9.i.l991, Nielsen & Edwards, Malaise #5 (1<3 1$, 
ANIC), 41.50S 146.03E, Pelion Hut 3km S Mt. Oakleigh, 850 m, 83-12. 
ii. 1991, A. Calder & W. Dressier, Malaise #5, closed forest (1 USNM). 
Description 
Male. Body length 3.3-3.5 mm. Anepisternal disc present. 
First flagellomere orbicular. Bristles black. Arista closely 
pubescent. Vibrissa relatively long and curved. Ocellar 
bristle long and relatively thin. Three dorsocentral bristles 
(decreasing in height anteriorly) with one small bristle 
in front of anterior dorsocentral. Gena high and sharply 
incurved medially at mid-height. Face and buccal cavity 
evenly curving and velvety. Frons dark brown with anterior 
margin orange, and lateral and posterior margins yellowish; 
antenna (excluding arista) yellow with first flagellomere dark 
brown excluding base; back of head dark brown (reddish 
above); occiput and clypeus dark brown; face yellow or with 
orange tint; remainder of head yellow; frons pilose to level 
of ocellar tubercle; dorsal half of gena silvery tomentose. 
Thorax dark brown. Fore legs dark brown with base of 
tibia yellow and fore coxa dirty yellow/brown and silvery 
tomentose with base brown; mid and hind legs dark brown 
with distal half of mid femur brownish to yellow, apex of 
coxae to base of femora yellow, mid tibia and tarsi light 
brown and hind tibia yellow with two dark bands. Abdomen 
dark brown. M 1+2 ratio approximately 6.0. Wing lightly 
clouded around cross veins and along anterodistal and distal 
margins. Halter white with stalk brownish (darker to base). 
Female. As described for male except as follows: first 
flagellomere infuscated on outer-dorsal margin, anterior 
margin and distal % of inner face; frons shiny laterally; 
tergites 2-5 slightly lighter. 
Male terminalia (Figs 85-87). Epandrium small and 
relatively wide. Cerci less than half height of epandrium and 
apically emarginate. Surstylus higher than long, rounded, 
almost bare on outer face and with inner-distal tubercle-like 
bristles. Hypandrium+pregonite triangular with two stout 
medial bristles and several setulae on elongate posteroventral 
Figures 88-106 [facing page]. Female internal genitalia; (88) Heteromeringia asteia n.sp.; (89) H. australiae Malloch; (90) H. australiae 
(aberrant NSW female); (91) H. australiae (abberant QLD female); (92) H. bisetosa n.sp.; (93) H. hardyi McAlpine; (94) H. helina n.sp.; 
(95) H. hypoleuca McAlpine; (96) H. imitans Malloch; (97) H. imitans (type 2 female); (98) H. imitans (type 3 female, CNC); (99) H. 
laticomis McAlpine; (100) H. limacens n.sp.; (101) H. macropa n.sp.; (102) H. patula n.sp.; (103) H. ptenopa n.sp.; (104) H. spinulosa 
McAlpine (dark gena, QLD); (105) H. spinulosa (pale gena, QLD); (106) H. trisetosa n.sp. 
