Editorial Board 
Dr Shane Ahyong 
Dr Don Colgan 
Dr Elena Kupriyanova 
Dr Andrew Mitchell 
Dr Robin Torrence 
Dr Ross Sadlier 
Editor 
Dr Shane McEvey 
Journal compilation © 2013 Australian Museum, Sydney 
No part of this publication may be reproduced without 
permission of the Editor. 
Volume 65 Number 3 
Published (print and online) 18 December 2013 
Price: AU$20.00 
Printed byRodenprint Pty Ltd, Sydney 
ISSN 0067-1975 (print) 
ISSN 2201-4349 (online) 
The Australian Museum is a statutory authority of, 
and principally funded by, the NSW State Government. 
NSW 
GOVERNMENT 
The Australian Museum houses some of the world’s most 
important collections of Australian animal, fossil and geological 
specimens and cultural objects. Research on these millions of 
specimens and artefacts yields insights into how our world changes 
through time and how its diversity can be classified, interpreted, 
and appreciated. This knowledge, when shared among the scientific 
and broader community—initially through publication—helps 
us understand human impact on our environment and what 
reasonable steps society can take now for the well-being of future 
generations. Our responsibility is to inspire the exploration of 
nature and cultures; our vision is a beautiful and sustainable 
natural world with vibrant and diverse cultures. 
Since 1889 the results of studies on Australian Museum 
collections, or studies that more generally lead to a better 
understanding of nature and cultures in the Australian Region, 
have been published by the Museum in its premier science 
journal Records of the Australian Museum. In 1999 we began 
releasing PDF of published articles through our open archive 
website. In 2008 we adopted DOI registration for our online 
content to facilitate permanence. In 2009 we digitized the entire 
legacy of all science published by us since 1851, and made that 
huge searchable resource permanently and freely available at our 
website. To accelerate publication of peer-reviewed science we are 
adopting a one article per publication model from volume 65 and 
we are limiting, but not abandoning, print production. All that is 
published in print is immediately and freely available online. 
Authors are invited to submit manuscripts to the Editor. 
Manuscripts meeting subject and stylistic requirements outlined 
in the Instructions to Authors are peer-reviewed by external referees 
to meet standards of excellence set by the Editorial Board. 
http://australianmuseum.net.au/ Scientific-Publications 
Records of the Australian Museum is covered in the Thomson 
Reuters Scientific services: Current Contents® / Agriculture, 
Biology, and Environmental Sciences, and Science Citation Index 
Expanded (also known as SciSearch®) 
We encourage cross-linking in the scientific literature by applying 
DOI registration to Australian Museum publications via CrossRef ® 
