60 
Records of the Australian Museum (2012) Vol. 64 
by lateral occipital suture (Fig. 27); male: surstylus somewhat 
elongate-falcate (Fig. 25) . driesseni, p. 67 
Supracervical cuticular dark zone less developed, not extending 
laterad of lateral occipital suture; male: surstylus otherwise. 8 
8 Proepimeral (stigmatal) bristle usually absent (check both sides); 
fore femur in posterior view usually showing rather diffuse dark 
greyish sub-basal posterodorsal zone and quite separate less diffuse 
dark subapical zone, on anterior surface sub-basal and distal 
brown zones usually separate; male: epandrium, in profile, without 
angular extension behind and above articulation with surstylus; 
surstylus as in Fig. 26 . macalpinei, p. 67 
-Proepimeral bristle usually present, small; fore femur usually 
without two such separate dark zones on posterior surface, on 
anterior surface sub-basal and distal brown zones touching or fused; 
male: epandrium, in profile, showing angular prominence behind 
articulation with surstylus; surstylus as in Fig. 24. obliqua, p. 66 
Helosciomyza fuscinevris (Macquart) 
Figs 16, 17 
Sciomyza fuscinevris Macquart, 1851: 276-277. Not 
homonym of Sciomyza fuscinervis Zetterstedt, 1838, now 
in Tetanocera Dumeril. 
Helosciomyza ferruginea Hendel, 1917: 33. N.syn. Not H. 
ferruginea : Steyskal, 1979:736, mixture of other species. 
Helosciomyza aliena Malloch, 1928: 324—325, synonymized 
D. McAlpine, 1992: 90. 
Helosciomyza fuscinevris (Macquart).—D. McAlpine, 
1992: 90. 
Type material. Holotype S of Sciomyza fuscinevris, 
Nouvelle-Hollande [New Holland = continental Australia, 
east coast added in publication—Macquart, 1851], anon. 
(PM, examined by D. McAlpine, 1990). Holotype S of 
Helosciomyza ferruginea, Victoria, 1888 (NMW, examined 
by D. McAlpine 1969, P. Sehnal 2008). Holotype S of 
Helosciomyza aliena. Broken Hill [western New South Wales], 
9.vi. 1925, anon. (AM K264571, formerly in SPHTM). 
Other material examined (localities only given). Northern 
Territory: Todd River, 9 km NE of Alice Springs (ANIC). 
Queensland: Eidsvold (ANIC). New South Wales: many specimens 
seen from many localities (not listed here), ranging from coast to 
far west of state and including higher ranges, mostly S of 32°S 
(AM, ANIC). Australian Capital Territory: numerous localities in 
Canberra district (ANIC). Victoria: Delegate River (AM); Benalla 
(AM); La Trobe University (AM, MV). Tasmania: Miena (ANIC); 
Lake Saint Clair (AM). South Australia: Mount Gambier (ANIC); 
Waite Park (ANIC); Mount Lofty (SAM); Ravine des Casoars, 
Kangaroo Island (AM); Stockport (ANIC); Maree (ANIC); Old 
Alton Downs, Simpson Desert (ANIC). Western Australia: Thomas 
River estuary, Esperance district (ANIC); Meerup Springs Farm, 
near Northcliffe (AM); near Mount Chudalup (AM); Frankland 
River (AM); Margaret River (AM). 
Distribution. Temperate Australia generally, approaching 
the tropics in Queensland and Northern Territory (near Alice 
Springs), south to Victoria, Tasmania, and southern coast 
of Western Australia; including some high-rainfall and arid 
areas, sea-level to moderate altitudes. 
Notes 
Helosciomyza fuscinevris is the only Australian species of the 
genus with pteropleural (anepimeral) setulae. Hypopleural 
setulae are usually also present, except in a few of the 
smallest specimens, and abdominal sternite 1 is setulose 
except in a small number of females, in which this sternite 
is more reduced than in males. 
According to Barnes (1981) this species is extremely 
similar to H. subalpina Tonnoir & Malloch, the only New 
Zealand species now retained in Helosciomyza. Barnes seems 
to infer that the c. 80 specimens of H. subalpina which he 
examined lack setulae on the ventral surface of vein 1 (or 
Rl). I have carefully examined 41 Australian specimens of H. 
fuscinevris, of which 40 possess one or more minute ventral 
setulae on vein 1. Also the dorsal striping of the thorax is 
less distinct in H. subalpina. These data seem to indicate 
a significant genetic difference between New Zealand and 
Australian populations, but this needs confirmation by more 
detailed morphological and molecular studies. 
The surstyli are slightly variable in shape, but in most 
specimens form an asymmetrical pair (Figs 16, 17); the 
cuticle on much of the posterior surface is thinner than 
elsewhere, almost membranous; pruinescence may be 
apparently absent or is present on part of the basal third. 
Helosciomyza bickeli n.sp. 
Figs 2, 15 
Helosciomyza ferruginea Hendel.-Steyskal & Knutson, 
1979:734-736 (misidentifications; fig. 16 probably refers 
to H. steyskali, fig. 18 refers to H. bickeli); Barnes, 1981: 
56-57 (in part, misidentification; fig. 13 does not represent 
H. bickeli). 
Type material. Holotype <$■ New South Wales: “Mooraback”, 
Werrikimbe National Park, 31°09'S 152°13'E, 980 m, 6.xii.l986, 
D.J.B. (AM K264288). Double-mounted on micro-pin through 
polyporus, postabdomen in micro-vial on pin. Paratypes. New South 
Wales: 1$, Gibraltar Range National Park, Feb. 1982,B.J.D. (AM); 
9SS, Mount Banda Banda, 1200 m, Dec. 1986, D.J.B. (AM, ANIC); 
\S, 2$9, Cobcroft Creek, Werrikimbe, Dec. 1986, D.J.B. (AM). 
Other material examined (localities only given). Queensland: 7 
miles (c. 11 km) up Mount Lewis Road, Julatten district (ANIC); 
Mount Edith, N of Tinaroo Dam (ANIC); Sluice Creek, near Millaa 
Millaa (ANIC); Paluma (AM); Eurimbula, Miriam Vale district 
(AM, QM); Summer Creek, near Kenilworth (AM); Mount Tenison 
Woods, near Mount Glorious (AM); Bundaroo Creek, Conondale 
Range (AM); Killarney (UQIC); Binna Burra, Lamington National 
Park (AM). New South Wales: Mount Warning, near Murwillumbah 
(AM); Falbrook, 30 km NNE of Singleton (AM); Barrington River, 
W of Gloucester (ANIC); Wallaroo State Forest, N of Raymond 
Terrace (ANIC); Myall Lakes (AM); Myall River (AM); Mount 
