70 
Records of the Australian Museum (2012) Vol. 64 
also the fore femur. The hind femur lacks the anterior bristle 
in both sexes, a condition approached in Neosciomyza (absent 
in male, reduced in female). The abdominal coloration is 
distinctive, the tergites being densely ashy grey-pruinescent 
with distinct dark brown zones. 
As previously indicated (D. Me Alpine, 1992), Cobergius 
includes a single known species. 
Cobergius vittatus (Macquart) 
Fig. 3 
Helomyza vittata Macquart, 1851: 279, pi. 25, fig. 6. 
Cobergius canus Barnes, 1981: 50-51, figs 1, 2. 
Cobergius hirsutus Barnes, 1981: 52. 
Cobergius vittatus (Macquart).-D. Me Alpine, 1992: 90-91. 
Type material. See list in D. Me Alpine (1992: 91). 
Other material examined. Victoria: 3 J $, Lady Julia Percy 
Island, W of Port Fairy, Jan. 1936, W.S. (AM, MV). 
Distribution. Kangaroo Island to Bass Strait Islands— 
apparently restricted to island shores. 
Notes 
Evidence for the above synonymy was discussed by D. 
Me Alpine (1992). Barnes stated, with regard to C. hirsutus , 
“The holotype female has two pairs of fine, black setae on 
the prosternum, hence the Latin name.” In fact, the holotype 
has one anterior setula on the left side of the prostemum only 
and two setulae on the membrane near the right side of the 
prosternum. Other specimens in which the prosternum is 
exposed have no setulae in this region. 
The possibility of an association with fur seal colonies 
is mentioned above. 
Acknowledgments. Nicole Fisher (ANIC), Ken Walker, and 
Catriona McPhee (MV) loaned significant collections under their 
care. Peter Sehnal (NMW) and Ted Edwards (ANIC) provided 
information on specimens. Stephen Marshall, Michael Driessen, and 
Gunther Theischinger provided important Tasmanian specimens. 
Suzanne Lindsay carried out electron microscopy. Dan Bickel, 
Shane McEvey, and Helen Smith made helpful comments on the 
manuscript. 
References 
Barnes, J. K. 1979. The taxonomic position of the New Zealand genus 
Prosochaeta Malloch (Diptera: Sciomyzidae). Proceedings of the 
Entomological Society of Washington 81: 285-297. 
Bames, J. K. 1980a. Biology and immature stages of Helosciomyza 
subalpina (Diptera: Helosciomyzidae), an ant-killing fly from 
New Zealand. New Zealand Journal of Zoology 7: 221-229. 
http://dx.doi.org/10.1080/03014223.1980.10423779 
Bames, J. K. 1980b. Immature stages of Polytocus costatus (Diptera: 
Helosciomyzidae) from the Snares Islands, New Zealand. New 
Zealand Journal of Zoology 7: 231-233. 
http://dx.doi.org/10.1080/03014223.1980.10423780 
Bames, J. K. 1981. Revision of the Helosciomyzidae (Diptera). 
Journal of the Royal Society of New Zealand 11: 45-72. 
http://dx.doi.org/10.1080/03036758.1981.10419451 
Bames, J. K. 1984. Biology and immature stages of Dryomyza anilis 
Fallen (Diptera: Dryomyzidae). Proceedings of the Entomological 
Society of Washington 86: 43-52. 
Burger, J. F., J. R. Anderson, and M. F. Knudsen. 1980. The habits 
and life history of Oedoparena glauca (Diptera: Dryomyzidae), a 
predator of barnacles. Proceedings of the Entomological Society 
ofWashington 82: 360-377. 
Colless, D. H., andD. K. McAlpine. 1991. Chapter 39. Diptera (flies). 
The Insects of Australia, second edition, pp. 717-786. Carlton: 
Melbourne University Press. 
Crosskey, R. W. 1973. A conspectus of the Tachinidae (Diptera) of 
Australia, including keys to the supraspecific taxa and taxonomic 
and host catalogues. Bulletin of the British Museum (Natural 
History) Entomology supplement 21: 221 pp. 
Griffiths, G. C. D. 1972. The phylogenetic classification of the Diptera 
Cyclorrhapha with special reference to the structure of the male 
postabdomen. 340 pp. The Hague: W. Junk. 
Hackman, W., and R. Vaisanen. 1985. The evolution and phylogenetic 
significance of the costal chaetotaxy in the Diptera. Annales 
Zoologici Eennici 22: 169-203. 
Harrison, R. A. 1959. Acalypterate Diptera of New Zealand. New 
Zealand Department of Scientific and Industrial Research, 
Bulletin 128: 382 + vii pp. 
Hendel, F. 1917. Beitrage zur Kenntis der acalyptraten Musciden. 
Deutsche Entomologische Zeitschrift 1917: 33-47. 
Macquart, P. J. M. 1851. Dipteres exotiques, suppl. 4 (second part): 
161-364, pis 15-28. 
Malloch, J. R. 1928. Notes on Australian Diptera, No. xv. Proceedings 
of the Linnean Society of New South Wales 53: 319-335. 
McAlpine, D. K. [1973]. The Australian Platystomatidae (Diptera, 
Schizophora) with a revision of five genera. Australian Museum 
Memoir 15: 1-256 (dated 1972, published 1973). 
http://dx.doi.org/10.3853/j.0067-1967.15.1972.454 
McAlpine, D. K. 1985. The Australian genera of Heleomyzidae 
(Diptera: Schizophora) and a reclassification of the family into 
tribes. Records of the Australian Museum 36: 203-251. 
http://dx.doi.org/10.3853/j.0067-1975.36.1985.346 
McAlpine, D. K. 1988. Studies in upside-down flies (Diptera: 
Neurochaetidae). Part II. Biology, adaptations, and specific mating 
mechanisms. Proceedings of the Linnean Society of New South 
Wales 110: 59-82. 
McAlpine, D. K. 1991a. Review of the Australian kelp flies (Diptera: 
Coelopidae). Systematic Entomology 16: 29-84. 
http://dx.doi.org/10.1111/j. 1365-3113.1991 ,tb00573.x 
McAlpine, D. K. 1991b. Relationships of the genus Heterocheila 
(Diptera: Sciomyzoidea) with description of a new family. 
Tijschrift voor Entomologie 134: 193-199. 
McAlpine, D. K. 1992. The earliest described species of Helo¬ 
sciomyzidae (Diptera: Schizophora). Australian Entomological 
Magazine 19: 89-92. 
McAlpine, D. K. 1995. Critical review of Australasian records of 
dryomyzid flies and resultant new combinations and synonymy in 
Lauxaniidae and Platystomatidae (Diptera: Schizophora). General 
and Applied Entomology 26: 41—43. 
McAlpine, D. K. 2007. The surge flies (Diptera: Canacidae: Zaleinae) 
of Australasia and notes on tethinid-canacid morphology and 
relationships. Records of the Australian Museum 59: 27-64. 
http://dx.doi.org/10.3853/j.0067-1975.59.2007.1468 
McAlpine, J. F. 1981. Chapter 2. Morphology and terminology — 
adults. In Manual ofNearctic Diptera 1, ed. J. F. McAlpine et al ., 
pp. 9-63. Hull, Quebec: Canadian Government Publishing Centre. 
Meier, R., and B. M. Wiegmann. 2002. A phylogenetic analysis of 
Coelopidae (Diptera) based on morphological and DNA sequence 
data. Molecular Phylogenetics and Evolution 25: 393^-07. 
http://dx.doi.org/10.1016/SI 055-7903(02)00276-2 
Ozerov, A. L. 1998. Family Dryomyzidae. Contributions to a Manual 
of Palaearctic Diptera 3: 349-355. Budapest: Science Herald. 
Steyskal, G. C. 1965. The subfamilies of the Sciomyzidae of the world 
(Diptera, Acalyptratae). Annals of the Entomological Society of 
America 58: 593-594. 
Steyskal, G. C. 1977. Family Dryomyzidae. A catalogue of the 
Diptera of the Americas south of the United States 62: 2 pp. 
Steyskal, G. C., and L. Knutson. [1979]. Helosciomyzinae in 
Australia (Diptera: Sciomyzidae). Australian Journal of Zoology 
26: 727-743. 
http://dx.doi.org/10.1071/ZO9780727 
