Zhen & Nicoll: Canning Basin Serratognathus bilobatus Fauna 
5 
Table 2. Comparison of occurrence of conodont species of the Serratognathus fauna from the Emanuel Formation with their 
distribution in other Serratognathus faunas recognized in China and other coeval successions referred to in the text. 
species 
a 
o 
O-I O 
S-H (J-h 
<D ^ 
Oh a) 
•i i 
B w 
o 
O 
gg e§ 
o W 
8 2 
U 3 
O 
O g 
S < 
<D 
o 
o 
N 
>3 
s 
£ 
o3 
O 
43 
u 
43 
o 
. o 
Co 00 
CD 
c 
o 
N 
si 
la 
-a 43 
■3 IS 
CO £ 
■s .a 
§> S3 
I « 
W> O 
O -o 
j§ 6 
N S3 
O 
S3 
3 
o | 
a £ 
Oh 
3S 
2 .a 
V a 
bapq 
5 a 
3 ' Ch 
S3 
o 
£ 
•a 
Q 
S3 
cd 
S3 
<D 
S3 
o 
C/3 
<D 
a 
j 
T3 
<D 
PQ 
CD 
S3 
o 
N 
(D 
Oh- 
in 
s 
X3 
S3 
S3 
S3 
O 
8 I 
03 Z 
Acodus deltatus? 
• • 
Acodus? transitans 
• 
Bergstroemognathus extensus 
• • 
• • • 
• • 
Cornuodus sp. 
• 
Drepanodus arcuatus 
• • 
• • 
• 
Drepanoistodus sp. cf. D. nowlani 
• 
• 
gen. et sp. indet. A 
• 
gen. et sp. indet. B 
• 
Fahraeusodus adentatus 
• 
Lissoepikodus nudus 
• • 
Nasusgnathus dolonus 
• • 
• • 
• 
Paracordylodus gracilis 
• 
• 
Paltodus sp. 
• 
Paroistodus parallelus 
• 
• • 
Paroistodus proteus 
• • 
• • 
• 
Prioniodus adami 
• •? 
• 
Protopanderodus gradatus 
• • 
• • 
• 
Protoprioniodus simplicissimus 
• • 
• 
• 
Scolopodus houlianzhaiensis 
• • 
• • • 
• 
Semiacontiodus sp. cf. S. cornuformis 
• 
• 
Serratognathus bilobatus 
• • 
• • • 
• • • 
Stiptognathus borealis 
• 
Triangulodus bifidus 
• • 
• 
• 
Tropodus australis 
• 
• 
Texas (Repetski, 1982), and the Argentine Precordillera 
(Fehnert, 1995; AlbanesiinAlbanesi etal., 1998). According 
this correlation gap. 
Triangulodus bifidus Zhen in Zhen et al., 2006 is a 
to Ethington et al. (2000), it occurs in the Fillmore Formation 
of Utah, in the upper quarter of the El Paso Group and in 
the upper member of the Marathon Fimestone of Texas, and 
in the Mountain Springs Formation of Nevada, constrained 
within an interval of upper Acodus deltatus/Oneotodus 
costatus Biozone to lower O. communis Biozone (early 
Floian age). However, in the Argentine Precordillera, earliest 
occurrence of S. borealis occurs in the upper P. proteus 
Biozone within the Tremadocian/Floian boundary interval. 
Scolopodus houlianzhaiensis An & Xu in An et al., 
1983 occurs abundantly in the Fiangjiashan Formation of 
North China with a similar stratigraphic range to that of 
Serratognathus bilobatus (An et al., 1983), and was also 
reported from the Honghuayuan Formation in Guizhou 
(South China) in association with Serratognathus diversus 
(Zhen et al., in press a). As both the Fiangjiashan Formation 
and Honghuayuan Formation were deposited in shallow 
subtidal settings, faunas recovered from these two units are 
dominated by endemic forms, and it is difficult to directly 
correlate them with either the Balto-Scandian or North 
American Mid-continent successions. Thus the presence 
of S. houlianzhaiensis in association with Serratognathus 
bilobatus in the Emanuel Formation is critical to bridging 
morphologically distinctive species occurring throughout 
the Honghuayuan Formation and also into the lower part 
of the overlying Meitan Formation in South China (Zhen 
et al., 2006, 2007a). It is apparently a rare species in the 
Emanuel Formation, represented by only a few specimens 
(Fig. 15I-K), and will be reported from the S. bilobatus 
fauna in the Arafura Basin of northern Australia (Nicoll 
unpublished material). 
Co-occurrence of Paroistodus proteus and P. parallelus in 
the Emanuel Formation also suggests a correlation with the 
upper part of the P. proteus Biozone of the Balto-Scandian 
succession. Fofgren (1997) indicated that in Sweden, 
Paroistodus parallelus makes its first appearance in the 
uppermost P. proteus Biozone, often associated with P. 
proteus, and extends to the upper middle O. evae Biozone. 
More specifically, occurrence of Prioniodus adami in the 
upper part of the Emanuel Formation allows a direct correla¬ 
tion with the P. adami Biozone (= upper P. proteus Biozone) 
recognized in the Cow Head Group of Newfoundland 
(Stouge & Bagnoli, 1988). There, P. adami was reported 
ranging through the lower to upper part (but not top) of Bed 
9 in the Fedge-Point of Head Section (Stouge & Bagnoli, 
1988, fig. 3), and the P. adami Biozone (succeeded by the P. 
